California Mixed Evergreen Forest

EVT 7043Mediterranean California Mixed Evergreen Forest
CES206.919GNRTreeConifer
Summary
This ecological system occurs from the Santa Cruz Mountains (and locally in the Santa Lucia Mountains), California, north into southwestern Oregon throughout the outer and middle Coast Ranges on Franciscan Formation soils (metasedimentary sandstones, schists, and shales) with moderate to high rainfall. This system occurs just inland from the redwood belt of this region. It also occurs in southern California in more mesic, protected, cooler sites of the Transverse and Peninsular ranges. Historic fire frequency in this system was higher than for redwood-dominated systems (every 50-100 years). It is characterized by mixes of coniferous and broad-leaved evergreen trees. Characteristic trees include Pseudotsuga menziesii, Quercus chrysolepis, Notholithocarpus densiflorus (= Lithocarpus densiflorus), Arbutus menziesii, Umbellularia californica, and Chrysolepis chrysophylla. On the eastern fringe of this system, in the western Siskiyous, other conifers occur such as Pinus ponderosa and Chamaecyparis lawsoniana. In southern California (Transverse and Peninsular ranges), Pseudotsuga macrocarpa replaces Pseudotsuga menziesii but co-occurs with Quercus chrysolepis and sometimes Quercus agrifolia. Calocedrus decurrens is occasional. In the southern portion of the range, Notholithocarpus densiflorus, Arbutus menziesii, Umbellularia californica, and Chrysolepis chrysophylla become less important or are absent. In the Santa Lucia Mountains, stands of Abies bracteata are included in this system and are an unusual and unique component. These stands are a mixture of Abies bracteata and Quercus chrysolepis. The more northerly stands tend to have dense or diverse shrub understories, with Corylus cornuta, Vaccinium ovatum, Rhododendron macrophyllum, Gaultheria shallon, Quercus sadleriana, Mahonia nervosa, and Toxicodendron diversilobum being common. Southern stands are less diverse and more sparse; Toxicodendron diversilobum is the most constant shrub, with Ribes spp. occasionally present, along with much Polystichum munitum. Especially in the south, stands are restricted to fire-protected sites (extremely steep, northerly, mesic slopes and coves) where fires from adjacent chaparral systems do not carry.
Source: NatureServe Explorer
Distribution
This system occurs from the Santa Lucia and Santa Cruz mountains of California north into southwestern Oregon throughout the outer and middle Coast Ranges and in southern California (Transverse and Peninsular ranges). It occurs in localized areas of the central to northern Sierra Nevada and southern and eastern Klamath Mountains.
Source: NatureServe Explorer
Ecologically Associated Plant Species

Plant species that characterize this ecosystem type, organized by vegetation stratum. These are species ecologically associated with the ecosystem, not confirmed present in any specific area.

Tree canopy

Abies bracteata, Arbutus menziesii, Calocedrus decurrens, Chamaecyparis lawsoniana, Pinus ponderosa, Pseudotsuga macrocarpa, Pseudotsuga menziesii, Quercus agrifolia, Quercus chrysolepis, Umbellularia californica

Shrub/sapling (tall & short)

Chrysolepis chrysophylla, Corylus cornuta, Lithocarpus densiflorus, Quercus sadleriana, Rhododendron macrophyllum

Short shrub/sapling

Berberis nervosa, Gaultheria shallon, Vaccinium ovatum

Herb (field)

Lilium occidentale, Polystichum munitum, Toxicodendron diversilobum

Nonvascular

Rubroboletus pulcherrimus
Source: NatureServe Ecological System assessment
Ecologically Associated Animals (15)

Animal species ecologically associated with this ecosystem type based on NatureServe assessment. These are species whose habitat requirements overlap with this ecosystem, not confirmed present in any specific roadless area.

Mammals (3)

Common NameScientific NameG-Rank
Dusky-footed WoodratNeotoma fuscipesG5
Pistol River Pocket GopherThomomys bottae detumidusG5T2Q
Gold Beach Pocket GopherThomomys mazama helleriG4T1T2

Birds (2)

Common NameScientific NameG-Rank
Oak TitmouseBaeolophus inornatusG4
Chestnut-backed ChickadeePoecile rufescensG5

Reptiles (1)

Common NameScientific NameG-Rank
Ring-necked SnakeDiadophis punctatusG5

Amphibians (2)

Common NameScientific NameG-Rank
California Slender SalamanderBatrachoseps attenuatusG5
EnsatinaEnsatina eschscholtziiG5

Butterflies & Moths (1)

Common NameScientific NameG-Rank
Mardon SkipperPolites mardonG2

Molluscs (5)

Common NameScientific NameG-Rank
Siskiyou ShoulderbandMonadenia chaceanaG2G3
Pacific SidebandMonadenia fidelis celeuthiaG4G5T2
Pacific SidebandMonadenia fidelis flavaG4G5T1T2
Scarletback Taildropper SlugProphysaon vanattaeG4G5
Siskiyou HesperianVespericola sierranusG3

Other (1)

Common NameScientific NameG-Rank
Umpqua SidebandMonadenia fidelis ssp. 2G4G5T1
Source: NatureServe Ecological System assessment
At-Risk Species Associated with this Ecosystem (11)

Species with conservation concern that are ecologically associated with this ecosystem type. G-Rank indicates global conservation status: G1 (critically imperiled) through G5 (secure). ESA status indicates U.S. Endangered Species Act listing.

Common NameScientific NameG-RankESA Status
Bristlecone FirAbies bracteataG2G3--
Western LilyLilium occidentaleG1G2Endangered
Siskiyou ShoulderbandMonadenia chaceanaG2G3--
Pacific SidebandMonadenia fidelis celeuthiaG4G5T2--
Pacific SidebandMonadenia fidelis flavaG4G5T1T2--
Umpqua SidebandMonadenia fidelis ssp. 2G4G5T1--
Mardon SkipperPolites mardonG2--
Red-pored BoleteRubroboletus pulcherrimusG3--
Pistol River Pocket GopherThomomys bottae detumidusG5T2Q--
Gold Beach Pocket GopherThomomys mazama helleriG4T1T2--
Siskiyou HesperianVespericola sierranusG3--
Source: NatureServe Ecological System assessment
Component Associations (5)

Plant community associations that occur within this ecological system. Associations are the finest level of the U.S. National Vegetation Classification (USNVC) and describe specific, repeating assemblages of plant species. Each association represents a distinct community type that may be found where this ecosystem occurs.

NameG-Rank
Chamaecyparis lawsoniana - Pseudotsuga menziesii / (Rhododendron macrophyllum) / Xerophyllum tenax ForestG1 NatureServe
Pseudotsuga menziesii / Quercus agrifolia ForestG3 NatureServe
Pseudotsuga menziesii - Quercus chrysolepis ForestG3 NatureServe
Pseudotsuga menziesii - (Umbellularia californica) / Frangula californica ForestG4 NatureServe
Pseudotsuga menziesii / Umbellularia californica / Polystichum munitum ForestG4 NatureServe
State Conservation Ranks (2)

Subnational conservation status ranks (S-ranks) assigned by Natural Heritage Programs in each state where this ecosystem occurs. S1 indicates critically imperiled at the state level, S2 imperiled, S3 vulnerable, S4 apparently secure, and S5 secure. An ecosystem may be globally secure but imperiled in specific states at the edge of its range.

StateS-Rank
CASNR
ORSNR
Roadless Areas (110)

Inventoried Roadless Areas where this ecosystem is present, identified from LANDFIRE 2024 Existing Vegetation Type spatial analysis. Coverage indicates the proportion of each area occupied by this ecosystem type.

California (85)

AreaForestCoverageHectares
Bakeoven RidgeShasta-Trinity National Forest79.2%15.12
Slide CreekSix Rivers National Forest78.7%3,645.99
Monkey CreekSix Rivers National Forest77.0%2,808.9
Orleans Mtn. BSix Rivers National Forest67.8%4,714.38
PacksaddleSix Rivers National Forest63.3%988.74
Cow CreekSix Rivers National Forest61.7%317.52
Redcap Addition Rare ISix Rivers National Forest61.6%121.77
UnderwoodSix Rivers National Forest60.8%1,620.99
Siskiyou BSix Rivers National Forest58.6%4,472.73
Ukonom CreekKlamath National Forest57.8%1,081.98
South ForkShasta-Trinity National Forest55.1%3,742.38
Blue Creek Rare ISix Rivers National Forest54.2%2,661.21
Cow CreekShasta-Trinity National Forest52.6%4,812.3
Bell QuinbyShasta-Trinity National Forest50.4%2,354.31
Ship MountainSix Rivers National Forest48.6%2,348.82
Fisher Gulch BShasta-Trinity National Forest47.8%927.09
China Springs BShasta-Trinity National Forest44.2%101.34
KellySix Rivers National Forest41.3%868.41
Weaver BallyShasta-Trinity National Forest39.2%132.03
PattisonShasta-Trinity National Forest39.0%4,625.1
PantherShasta-Trinity National Forest38.2%1,857.42
SiskiyouKlamath National Forest37.6%8,218.26
Dog CreekShasta-Trinity National Forest37.4%756.54
PortugueseKlamath National Forest33.7%2,576.43
Orleans Mtn.Klamath National Forest30.4%6,042.51
Kangaroo (CA)Rogue River National Forests30.2%728.46
Hobo GulchShasta-Trinity National Forest29.6%10.53
Little French CShasta-Trinity National Forest29.5%1,377.36
BlackKlamath National Forest29.2%772.38
ChinquapinShasta-Trinity National Forest28.0%2,501.64
UnderwoodShasta-Trinity National Forest27.8%342.27
Kettle Mtn.Shasta-Trinity National Forest26.7%495.72
West GirardShasta-Trinity National Forest26.2%3,973.95
East GirardShasta-Trinity National Forest25.8%2,914.47
Wells MountainShasta-Trinity National Forest25.6%612.81
KangarooKlamath National Forest24.6%4,035.78
Board CampSix Rivers National Forest23.9%436.95
SoliderSix Rivers National Forest23.4%1,412.46
Orleans Mtn. CSix Rivers National Forest21.8%1,374.3
MuseKlamath National Forest21.2%20.52
Mt. LassicSix Rivers National Forest19.5%523.89
Stoveleg GapShasta-Trinity National Forest19.0%4.68
Siskiyou ASix Rivers National Forest18.7%76.86
JohnsonKlamath National Forest17.5%753.12
North Fork SmithSix Rivers National Forest17.4%2,666.16
Pilot CreekSix Rivers National Forest17.4%645.75
ChanchelullaShasta-Trinity National Forest17.0%269.28
BackboneShasta-Trinity National Forest16.3%758.43
EagleShasta-Trinity National Forest15.2%404.19
South KalmiopsisSix Rivers National Forest15.1%19.62
Devils RockShasta-Trinity National Forest15.1%990.72
Salt GulchShasta-Trinity National Forest14.7%386.1
Jacobs (fs)Klamath National Forest14.1%32.85
Flem (FS)Klamath National Forest13.6%31.23
GriderKlamath National Forest13.2%568.62
Condrey MountainRogue River National Forests10.1%369.81
West BeegumShasta-Trinity National Forest9.4%198.18
ThatcherMendocino National Forest8.9%598.86
East BeegumShasta-Trinity National Forest8.6%292.14
Salt CreekSix Rivers National Forest8.4%324.9
KelseyKlamath National Forest7.1%92.61
Thomes CreekMendocino National Forest6.1%411.75
Skeleton GladeMendocino National Forest6.0%225.9
SiskiyouSiskiyou National Forests6.0%73.8
Tom MartinKlamath National Forest5.1%188.1
Snow MountainMendocino National Forest4.7%272.52
East ForkShasta-Trinity National Forest4.6%114.39
Castle Crags BShasta-Trinity National Forest4.5%29.43
Elk CreekMendocino National Forest4.4%416.43
SnoozerKlamath National Forest4.1%390.78
BoulderKlamath National Forest3.8%6.66
BriscoeMendocino National Forest3.8%109.71
Slate CreekShasta-Trinity National Forest3.7%99
Reister CanyonMendocino National Forest3.6%86.76
Big Butte ShinboneSix Rivers National Forest3.3%14.85
Bonanza KingShasta-Trinity National Forest3.3%218.43
Black ButteMendocino National Forest3.3%205.65
Penney RidgeShasta-Trinity National Forest3.2%67.05
CrapoKlamath National Forest2.9%17.64
Indian CreekKlamath National Forest2.7%55.62
GrindstoneMendocino National Forest2.4%252.54
Chalk PeakLos Padres National Forest1.6%48.96
RussianKlamath National Forest1.6%141.84
Bear CanyonLos Padres National Forest1.1%63.63
Deer MountainMendocino National Forest0.8%38.52

Oregon (25)

AreaForestCoverageHectares
Mt. EmilySiskiyou National Forests94.8%2,210.31
QuosatanaSiskiyou National Forests93.5%1,945.44
Potato MountainSiskiyou National Forests91.9%2,214.99
RogueSiskiyou National Forests90.7%1,237.23
Shasta CostaSiskiyou National Forests78.7%4,593.15
Kalmiopsis AdditionSiskiyou National Forests78.3%413.64
BriggsSiskiyou National Forests75.0%1,795.5
Windy ValleySiskiyou National Forests71.1%2,851.92
Mule CreekSiskiyou National Forests64.7%106.29
PacksaddleSiskiyou National Forests64.5%1,440.72
North KalmiopsisSiskiyou National Forests58.1%21,541.59
Copper MountainSiskiyou National Forests58.1%2,065.59
Grassy KnobSiskiyou National Forests50.2%1,150.92
KinneyRogue River National Forests44.1%1,339.65
Nihwik MountainSiskiyou National Forests37.9%1,211.85
North Fork SmithSiskiyou National Forests35.4%131.13
South KalmiopsisSiskiyou National Forests29.5%12,468.6
Little GraybackRogue River National Forests22.1%713.25
Kangaroo (OR)Rogue River National Forests12.9%753.57
Bitter LickRogue River National Forests1.8%48.69
KangarooSiskiyou National Forests1.3%37.44
Brown MountainRogue River National Forests1.3%34.92
Mcdonald PeakRogue River National Forests1.3%48.87
DoneganUmpqua National Forest1.2%28.89
LastUmpqua National Forest1.1%33.93
Methodology and Data Sources

Ecosystem classification: Ecosystems are classified using the LANDFIRE 2024 Existing Vegetation Type (EVT) layer, mapped to NatureServe Terrestrial Ecological Systems via a curated crosswalk. Each EVT is linked to the USNVC (U.S. National Vegetation Classification) hierarchy through pixel-level co-occurrence analysis of LANDFIRE EVT and NatureServe IVC Group rasters across all roadless areas.

Vegetation coverage: Coverage percentages and hectares are derived from zonal statistics of the LANDFIRE 2024 EVT raster intersected with roadless area boundaries.

Ecosystem narratives and community species: Sourced from the NatureServe Explorer API, representing professional ecological assessments of vegetation composition, environmental setting, dynamics, threats, and characteristic species assemblages.

IVC hierarchy: The International Vegetation Classification hierarchy is sourced from the USNVC v3.0 Catalog, providing the full classification from Biome through Association levels.

Component associations: Plant community associations listed as components of each NatureServe Ecological System. Association data from the NatureServe Explorer API.

State ranks: Conservation status ranks assigned by NatureServe member programs in each state where the ecosystem occurs.