California Mountain Chaparral

EVT 7098California Montane Woodland and Chaparral
CES206.925GNRShrubShrubland
Summary
This ecological system includes chaparral or open shrubby woodlands found among montane forests above 1500 m (4550 feet) elevation from the southern Cascades of Oregon to the Peninsular Ranges of California into Baja California, Mexico, where much annual precipitation occurs as snow. These are often locations with steep, exposed slopes with rocky and/or shallow soils, often glaciated. Stands are not found in the foothills but rather occur commonly above 1524 m (5000 feet) in elevation. These are mosaics of woodlands with chaparral understories, shrub-dominated chaparral, or short-lived chaparral with conifer species invading, if good seed source is available. Shrubs will often have higher densities than the trees, which are more limited due to the rocky/thin soils. These can also be short-duration chaparrals in previously forested areas that have experienced crown fires. Trees tend to have a scattered open canopy or can be clustered, over a usually continuous dense shrub layer. Trees can include Pinus jeffreyi, Abies lowiana (= Abies concolor var. lowiana), Abies magnifica, Pinus monticola, Pinus lambertiana, Pinus coulteri, Pinus attenuata, Hesperocyparis forbesii (= Cupressus forbesii), Hesperocyparis stephensonii (= Cupressus arizonica ssp. stephensonii), and Hesperocyparis nevadensis (= Cupressus nevadensis). Typical sclerophyllous chaparral shrubs include Arctostaphylos nevadensis, Arctostaphylos patula, Arctostaphylos glandulosa, Ceanothus cordulatus, Ceanothus diversifolius, Ceanothus pinetorum, Ceanothus velutinus, and Chrysolepis sempervirens (= Castanopsis sempervirens). Some stands can be dominated by winter deciduous shrubs, such as Prunus emarginata, Prunus subcordata and Ceanothus sanguineus (in Oregon), Prunus virginiana, Ceanothus integerrimus, Holodiscus discolor (= Holodiscus microphyllus), and Quercus garryana var. fruticosa (= var. breweri). Most chaparral species are fire-adapted, resprouting vigorously after burning or producing fire-resistant seeds. Occurrences of this system likely shift across montane forested landscapes with catastrophic fire events.
Source: NatureServe Explorer
Environment
[from M094] These are chaparral or open shrublands found at montane elevations throughout much of the western U.S., from the Sierra Nevada and Cascades and into the western Great Basin, Colorado Plateau, and Rocky Mountains. They occur in summer-dry habitats from 800 to 3000 m elevation. Can occur as low as 50 m in California, but mostly is found above 1500 m. Much of the precipitation comes as winter snow, and summer drought-stress is characteristic. These shrublands are mostly found on steep, usually south-facing or exposed slopes, where soils are rocky, shallow and well-drained, often glaciated. These are typically zonal disclimax or, occasionally, edaphic climax brushfields which occur in association with dry needle-leaved evergreen forests or woodlands. These shrublands are typically established after stand-replacing fires or clearcut logging in montane conifer forests or pinyon-juniper woodlands, and may be seral to forest after several decades. Excessively rocky or droughty, fire-prone sites in the forest may support relatively persistent stands of this macrogroup. These are in mosaics of woodlands and chaparral and may have conifer species invading if good seed source is available.
Source: NatureServe Explorer
Dynamics
[from M094] Two phases are recognized: first, early-seral and post-fire or post-logging shrub fields with few conifers; and second, edaphically controlled sites, with soils that are too dry or shallow-soiled for trees, hence sites where shrubs stay dominant (such as Quercus vacciniifolia, Chrysolepis sempervirens). Most chaparral species are fire-adapted, resprouting vigorously after burning or producing fire-resistant seeds. Occurrences of this macrogroup likely shift across montane forested landscapes with catastrophic fire events. Clearcut logging can also trigger regeneration of some of the chaparral species.
Source: NatureServe Explorer
Distribution
This system occurs above 1500 m (4550 feet) elevation from the southern Cascades of Oregon to the Klamath Mountains and Peninsular Ranges of California into Baja California, Mexico.
Source: NatureServe Explorer
Ecologically Associated Plant Species

Plant species that characterize this ecosystem type, organized by vegetation stratum. These are species ecologically associated with the ecosystem, not confirmed present in any specific area.

Tree canopy

Abies concolor var. lowiana, Abies magnifica, Hesperocyparis forbesii, Hesperocyparis nevadensis, Hesperocyparis stephensonii, Pinus attenuata, Pinus coulteri, Pinus jeffreyi, Pinus lambertiana, Pinus monticola

Shrub/sapling (tall & short)

Arctostaphylos glandulosa, Arctostaphylos nevadensis, Arctostaphylos patula, Ceanothus cordulatus, Ceanothus diversifolius, Ceanothus integerrimus, Ceanothus pinetorum, Ceanothus sanguineus, Ceanothus velutinus, Chrysolepis sempervirens, Holodiscus discolor, Prunus emarginata, Prunus subcordata, Prunus virginiana, Quercus garryana var. breweri

Herb (field)

Lilium occidentale, Limnanthes floccosa ssp. bellingeriana, Plagiobothrys figuratus ssp. corallicarpus
Source: NatureServe Ecological System assessment
Ecologically Associated Animals (7)

Animal species ecologically associated with this ecosystem type based on NatureServe assessment. These are species whose habitat requirements overlap with this ecosystem, not confirmed present in any specific roadless area.

Reptiles (1)

Common NameScientific NameG-Rank
GophersnakePituophis cateniferG5

Butterflies & Moths (1)

Common NameScientific NameG-Rank
Mardon SkipperPolites mardonG2

Insects (1)

Common NameScientific NameG-Rank
Siskiyou Slant-face GrasshopperChloealtis aspasmaG3

Molluscs (4)

Common NameScientific NameG-Rank
Shasta PebblesnailFluminicola multifariusG2
Siskiyou ShoulderbandMonadenia chaceanaG2G3
Pacific SidebandMonadenia fidelis flavaG4G5T1T2
Scaly ChaparralTrilobopsis loricataG2G3
Source: NatureServe Ecological System assessment
At-Risk Species Associated with this Ecosystem (15)

Species with conservation concern that are ecologically associated with this ecosystem type. G-Rank indicates global conservation status: G1 (critically imperiled) through G5 (secure). ESA status indicates U.S. Endangered Species Act listing.

Common NameScientific NameG-RankESA Status
Trailing WhitethornCeanothus diversifoliusG3?--
Coville's CeanothusCeanothus pinetorumG3--
Siskiyou Slant-face GrasshopperChloealtis aspasmaG3--
Shasta PebblesnailFluminicola multifariusG2--
Tecate CypressHesperocyparis forbesiiG2--
Paiute CypressHesperocyparis nevadensisG2--
Cuyamaca CypressHesperocyparis stephensoniiG1--
Western LilyLilium occidentaleG1G2Endangered
Bellinger's MeadowfoamLimnanthes floccosa ssp. bellingerianaG4T3--
Siskiyou ShoulderbandMonadenia chaceanaG2G3--
Pacific SidebandMonadenia fidelis flavaG4G5T1T2--
Coral-seeded AllocaryaPlagiobothrys figuratus ssp. corallicarpusG4T1T2--
Mardon SkipperPolites mardonG2--
Brewer OakQuercus garryana var. breweriG5T3--
Scaly ChaparralTrilobopsis loricataG2G3--
Source: NatureServe Ecological System assessment
Component Associations (9)

Plant community associations that occur within this ecological system. Associations are the finest level of the U.S. National Vegetation Classification (USNVC) and describe specific, repeating assemblages of plant species. Each association represents a distinct community type that may be found where this ecosystem occurs.

NameG-Rank
Arctostaphylos patula Sierran Chaparral ShrublandG5 NatureServe
Arctostaphylos viscida Sierran Chaparral ShrublandG5 NatureServe
Ceanothus cordulatus ShrublandG3 NatureServe
Ceanothus cordulatus / Sparse Understory Sierran ShrublandG4 NatureServe
Chrysolepis sempervirens ShrublandG4 NatureServe
Chrysolepis sempervirens / Sparse Understory Sierran ShrublandG3 NatureServe
Holodiscus discolor - Sambucus racemosa ShrublandGNR NatureServe
Holodiscus discolor / Sedum obtusatum ssp. boreale - Cryptogramma acrostichoides ShrublandGNR NatureServe
Prunus emarginata Sierran Chaparral ShrublandG3 NatureServe
State Conservation Ranks (2)

Subnational conservation status ranks (S-ranks) assigned by Natural Heritage Programs in each state where this ecosystem occurs. S1 indicates critically imperiled at the state level, S2 imperiled, S3 vulnerable, S4 apparently secure, and S5 secure. An ecosystem may be globally secure but imperiled in specific states at the edge of its range.

StateS-Rank
CASNR
ORSNR
Roadless Areas (241)

Inventoried Roadless Areas where this ecosystem is present, identified from LANDFIRE 2024 Existing Vegetation Type spatial analysis. Coverage indicates the proportion of each area occupied by this ecosystem type.

California (211)

AreaForestCoverageHectares
Raywood Flat ASan Bernardino National Forest83.6%13.77
Mokelumne - PleasantHumboldt-Toiyabe National Forest82.9%90.81
Mokelumne - SprattHumboldt-Toiyabe National Forest70.0%173.61
Big Butte ShinboneSix Rivers National Forest61.8%275.49
Indian CreekKlamath National Forest57.1%1,158.75
Salt CreekSix Rivers National Forest54.4%2,092.14
Iceberg - RodriquezHumboldt-Toiyabe National Forest53.1%49.14
Damon ButteModoc National Forest51.4%5,202.36
Adams PeakPlumas National Forest49.4%1,056.87
Wilderness ContiguousMendocino National Forest48.6%709.11
Waterhouse AHumboldt-Toiyabe National Forest48.0%6.39
Big Butte ShinboneMendocino National Forest46.8%807.21
ShuteyeSierra National Forest44.4%1,313.91
SoliderSix Rivers National Forest42.1%2,542.68
San JoaquinSierra National Forest41.7%3,790.98
Silver HillHumboldt-Toiyabe National Forest39.8%712.17
Slate CreekShasta-Trinity National Forest39.3%1,055.25
SugarloafSan Bernardino National Forest39.0%1,294.02
Chips CreekLassen National Forest38.0%4,474.98
Barney RileyHumboldt-Toiyabe National Forest35.2%1,150.83
Black ButteMendocino National Forest34.1%2,132.55
WoodchuckSierra National Forest33.9%36.81
Mokelumne - Hawkins PkHumboldt-Toiyabe National Forest33.8%2,259.54
Caples CreekEldorado National Forest33.1%2,391.48
Elk CreekMendocino National Forest32.2%3,021.03
Mokelumne - Mt. BullionHumboldt-Toiyabe National Forest31.9%1,755.36
Robinson PeakHumboldt-Toiyabe National Forest31.8%751.86
Hoover - EastHumboldt-Toiyabe National Forest31.8%11.88
Raywood Flat BSan Bernardino National Forest29.7%1,367.28
Domeland Add.Sequoia National Forest29.0%357.03
EbbettsHumboldt-Toiyabe National Forest27.9%28.8
Iceberg - Mill CreekHumboldt-Toiyabe National Forest27.7%3,025.35
Penney RidgeShasta-Trinity National Forest27.4%579.06
Jobs Peak (CA)Humboldt-Toiyabe National Forest27.3%2,496.96
Cherry LakeStanislaus National Forest27.1%121.23
EagleShasta-Trinity National Forest27.1%718.74
Bald RockPlumas National Forest27.1%512.37
SnoozerKlamath National Forest25.9%2,452.5
Flem (FS)Klamath National Forest25.6%58.68
Salt SpringsEldorado National Forest25.5%13.77
WaterhouseStanislaus National Forest25.4%451.98
Dinkey LakesSierra National Forest24.9%3,444.03
MayfieldLassen National Forest23.3%1,361.43
Bald MountainTahoe National Forest22.6%533.52
RinconSequoia National Forest22.5%4,983.03
Raymond PeakStanislaus National Forest22.5%332.64
Rose - IcelandHumboldt-Toiyabe National Forest22.5%106.47
PyramidEldorado National Forest21.8%2,146.95
Bell MeadowStanislaus National Forest21.5%693.63
Slate Mtn.Sequoia National Forest21.5%1,068.84
DardanellesEldorado National Forest21.4%702.9
Heartbreak RidgeSan Bernardino National Forest21.3%384.39
Castle PeakTahoe National Forest21.3%1,288.53
Granite Chief NLake Tahoe Basin Management Unit21.2%50.76
Mokelumne - CharityHumboldt-Toiyabe National Forest21.1%17.19
Wild Cattle MtnLassen National Forest20.5%412.56
ThatcherMendocino National Forest20.4%1,376.55
Dennison PeakSequoia National Forest20.0%508.32
Grizzly PeakPlumas National Forest19.6%494.19
Mt. RaymondSierra National Forest19.6%552.87
LavaLassen National Forest19.1%462.24
Chips CreekPlumas National Forest18.5%968.31
Carson - IcebergStanislaus National Forest18.4%4,191.12
Granite Chief SLake Tahoe Basin Management Unit18.1%41.49
Hoover - NorthHumboldt-Toiyabe National Forest18.1%115.11
KangarooKlamath National Forest17.3%2,848.5
Horse Mdw.Inyo National Forest16.9%389.52
Condrey Mtn.Klamath National Forest16.6%196.11
SiskiyouSiskiyou National Forests16.3%200.07
Tragedy - Elephants BackEldorado National Forest16.3%1,375.83
MosesSequoia National Forest16.3%1,454.85
Mt. Shasta BShasta-Trinity National Forest16.0%181.62
Mystic (CA)Humboldt-Toiyabe National Forest15.8%53.46
JohnsonKlamath National Forest15.5%670.05
EagleStanislaus National Forest15.3%997.02
Iceberg - SlinkardHumboldt-Toiyabe National Forest15.2%95.49
Fawn LakeEldorado National Forest15.0%70.11
ChinquapinShasta-Trinity National Forest14.9%1,325.34
Granite ChiefTahoe National Forest14.8%391.77
Middle ForkPlumas National Forest14.1%1,668.96
Hoover - Emma LakeHumboldt-Toiyabe National Forest14.0%57.33
Orleans Mtn. CSix Rivers National Forest14.0%880.56
Big CanyonModoc National Forest14.0%373.41
Keddie RidgeLassen National Forest13.9%217.35
Mt. HarknessLassen National Forest13.5%10.71
FreelLake Tahoe Basin Management Unit13.4%833.67
DomeStanislaus National Forest13.3%594.81
Bucks LakePlumas National Forest13.2%36.36
GriderKlamath National Forest13.2%567.45
Skeleton GladeMendocino National Forest13.2%491.58
Hoover - Cattle CkHumboldt-Toiyabe National Forest13.0%31.41
Grouse LakesTahoe National Forest12.9%998.28
Mt. EddyShasta-Trinity National Forest12.7%371.43
WoodpeckerSequoia National Forest12.7%612.27
PantherShasta-Trinity National Forest12.7%615.78
Monkey CreekSix Rivers National Forest12.7%461.7
Lion RidgeSequoia National Forest12.5%266.67
Raymond PeakEldorado National Forest12.5%127.35
Pleasant ViewAngeles National Forest12.1%1,294.38
Mt. RebaStanislaus National Forest12.0%187.65
ChicoSequoia National Forest11.9%1,922.4
Duncan CanyonTahoe National Forest11.7%408.06
DardanellesLake Tahoe Basin Management Unit11.2%645.21
Bonanza KingShasta-Trinity National Forest11.1%738.9
Mt. OlsenInyo National Forest11.0%96.48
Tom MartinKlamath National Forest11.0%403.29
Poison HoleEldorado National Forest10.7%114.12
RussianKlamath National Forest10.7%943.02
Lakes BasinTahoe National Forest9.8%22.14
KelseyKlamath National Forest9.8%128.61
Jennie LakeSequoia National Forest9.6%92.97
WoolstaffSequoia National Forest9.4%1,579.14
PyramidLake Tahoe Basin Management Unit9.3%292.32
Independence CreekInyo National Forest9.2%501.84
ChannellSequoia National Forest9.2%1,690.47
CrapoKlamath National Forest8.9%53.46
DryModoc National Forest8.7%272.88
SiskiyouKlamath National Forest8.7%1,906.2
Table Mtn.Inyo National Forest8.5%144.9
South ForkShasta-Trinity National Forest7.9%539.1
Buckeye RidgeHumboldt-Toiyabe National Forest7.7%69.48
Cub (fs)Klamath National Forest7.6%4.14
West BeegumShasta-Trinity National Forest7.6%158.85
ShacklefordKlamath National Forest7.5%50.58
San JoaquinInyo National Forest7.3%583.11
Sheep MountainAngeles National Forest7.3%625.59
Heart LakeLassen National Forest7.3%276.12
Hoover - Green Ck NoHumboldt-Toiyabe National Forest7.3%210.78
PattisonShasta-Trinity National Forest7.2%855.72
UnderwoodShasta-Trinity National Forest7.1%87.66
East BeegumShasta-Trinity National Forest7.0%240.21
Ukonom CreekKlamath National Forest7.0%131.49
Snow MountainMendocino National Forest7.0%409.41
Lake EleanorShasta-Trinity National Forest6.9%11.07
Butt Mtn.Lassen National Forest6.9%229.68
Thomes CreekMendocino National Forest6.9%463.5
Orleans Mtn.Klamath National Forest6.8%1,353.15
SherwinInyo National Forest6.3%80.01
Knox Mtn.Modoc National Forest6.2%149.85
Wonoga Pk.Inyo National Forest6.1%277.92
Pilot CreekSix Rivers National Forest6.0%224.73
North LakeInyo National Forest6.0%58.77
Hoover - Rickey PkHumboldt-Toiyabe National Forest6.0%329.13
South SierraSequoia National Forest5.9%190.53
Pyramid Peak BSan Bernardino National Forest5.8%168.48
AgnewSequoia National Forest5.8%222.75
Devils GardenLassen National Forest5.7%77.31
Trumbull PeakStanislaus National Forest5.6%138.69
West GirardShasta-Trinity National Forest5.5%832.5
North Fork Middle Fork American RiverTahoe National Forest5.3%242.73
Little French CShasta-Trinity National Forest5.2%243.36
Kangaroo (CA)Rogue River National Forests5.2%125.82
BackboneShasta-Trinity National Forest4.9%228.24
West YubaPlumas National Forest4.9%120.42
Log Cabin SaddlebagInyo National Forest4.7%291.06
South SierraInyo National Forest4.6%781.92
Laurel McgeeInyo National Forest4.5%159.84
PortugueseKlamath National Forest4.5%342.9
East YubaTahoe National Forest4.4%320.85
Onion SpringsLassen National Forest4.4%42.75
Jacobs (fs)Klamath National Forest4.4%10.17
Devil GulchSierra National Forest4.3%532.71
Cow CreekShasta-Trinity National Forest4.2%389.07
BriscoeMendocino National Forest4.1%120.51
ProspectLassen National Forest4.0%66.69
Rock Creek WestInyo National Forest3.8%54.99
Kings RiverSierra National Forest3.7%786.6
North Fork American RiverTahoe National Forest3.7%568.8
Sespe - FrazierLos Padres National Forest3.6%1,542.51
East GirardShasta-Trinity National Forest3.5%399.24
North MountainStanislaus National Forest3.5%109.71
Fisher Gulch BShasta-Trinity National Forest3.4%66.51
East ForkShasta-Trinity National Forest3.4%85.05
NightStanislaus National Forest3.3%42.93
Black Mtn.Sequoia National Forest3.2%194.76
AntimonyLos Padres National Forest3.0%497.79
ChanchelullaShasta-Trinity National Forest3.0%46.8
CypressLassen National Forest2.9%39.24
Box CampKlamath National Forest2.8%9.54
GrindstoneMendocino National Forest2.7%289.44
Mt. LassicSix Rivers National Forest2.7%72.09
West YubaTahoe National Forest2.6%171.63
Pyramid Peak ASan Bernardino National Forest2.5%145.98
Bell QuinbyShasta-Trinity National Forest2.5%116.28
Timbered CraterLassen National Forest2.4%40.59
Castle Crags BShasta-Trinity National Forest2.3%15.39
Snow MountainLassen National Forest2.2%19.17
TinemahaInyo National Forest2.1%229.23
Mill PeakSan Bernardino National Forest2.0%64.71
Cub CreekLassen National Forest1.9%64.8
Condrey MountainRogue River National Forests1.8%67.68
Coyote SoutheastInyo National Forest1.8%391.14
Reister CanyonMendocino National Forest1.8%43.38
Greenhorn CreekSequoia National Forest1.8%202.86
Ship MountainSix Rivers National Forest1.7%83.88
Wells MountainShasta-Trinity National Forest1.6%39.06
Orleans Mtn. BSix Rivers National Forest1.6%112.41
Hoover - Virginia LksHumboldt-Toiyabe National Forest1.4%28.35
Coyote NorthInyo National Forest1.4%65.7
Mill CreekSequoia National Forest1.4%152.01
PowleyModoc National Forest1.2%29.7
Cucamonga BSan Bernardino National Forest1.2%56.16
RubiconEldorado National Forest1.1%22.23
Sawmill - BadlandsLos Padres National Forest1.0%216.81
Glass MountainInyo National Forest1.0%221.76
Middle YubaTahoe National Forest1.0%29.88
Deep CreekSan Bernardino National Forest1.0%94.77
Hat Mtn.Modoc National Forest0.9%35.01
Siskiyou BSix Rivers National Forest0.9%66.69
Sears FlatModoc National Forest0.7%38.25
North Fork SmithSix Rivers National Forest0.4%64.44

Nevada (17)

AreaForestCoverageHectares
Rose - EvansHumboldt-Toiyabe National Forest41.8%810.09
Rose - Gray CreekHumboldt-Toiyabe National Forest38.1%56.25
Jobs Peak (NV)Humboldt-Toiyabe National Forest37.1%201.51
Rose - Alum CreekHumboldt-Toiyabe National Forest32.5%112.14
Rose - Dutch LouieHumboldt-Toiyabe National Forest30.3%44.73
Rose - Whites CanyonHumboldt-Toiyabe National Forest27.3%283.23
Rose - VerdiHumboldt-Toiyabe National Forest26.3%122.67
Rose - GalenaHumboldt-Toiyabe National Forest24.8%372.06
Rose - Thomas Mdw.Humboldt-Toiyabe National Forest23.3%33.93
Mt. Rose RoadlessLake Tahoe Basin Management Unit16.1%34.29
Rose - North NotchHumboldt-Toiyabe National Forest16.0%9.09
Rose - NortheastHumboldt-Toiyabe National Forest14.1%31.23
LincolnLake Tahoe Basin Management Unit12.8%340.92
Rose - Davis Mdw.Humboldt-Toiyabe National Forest10.9%103.86
Mystic (NV)Humboldt-Toiyabe National Forest8.3%189.45
Rose - BroncoHumboldt-Toiyabe National Forest8.3%28.98
Rose - Hunter Lk NoHumboldt-Toiyabe National Forest8.1%4.86

Oregon (13)

AreaForestCoverageHectares
Little GraybackRogue River National Forests6.5%211.23
Brattain ButteFremont National Forest5.5%133.83
Windy ValleySiskiyou National Forests5.1%203.22
Nihwik MountainSiskiyou National Forests4.9%157.77
North Fork SmithSiskiyou National Forests3.4%12.78
North KalmiopsisSiskiyou National Forests3.3%1,219.95
Kangaroo (OR)Rogue River National Forests2.7%155.97
South KalmiopsisSiskiyou National Forests2.0%865.17
Mcdonald PeakRogue River National Forests1.5%58.05
KangarooSiskiyou National Forests1.4%40.14
KinneyRogue River National Forests1.3%39.96
BriggsSiskiyou National Forests1.2%27.9
Crane MountainFremont National Forest0.9%85.05
Methodology and Data Sources

Ecosystem classification: Ecosystems are classified using the LANDFIRE 2024 Existing Vegetation Type (EVT) layer, mapped to NatureServe Terrestrial Ecological Systems via a curated crosswalk. Each EVT is linked to the USNVC (U.S. National Vegetation Classification) hierarchy through pixel-level co-occurrence analysis of LANDFIRE EVT and NatureServe IVC Group rasters across all roadless areas.

Vegetation coverage: Coverage percentages and hectares are derived from zonal statistics of the LANDFIRE 2024 EVT raster intersected with roadless area boundaries.

Ecosystem narratives and community species: Sourced from the NatureServe Explorer API, representing professional ecological assessments of vegetation composition, environmental setting, dynamics, threats, and characteristic species assemblages.

IVC hierarchy: The International Vegetation Classification hierarchy is sourced from the USNVC v3.0 Catalog, providing the full classification from Biome through Association levels.

Component associations: Plant community associations listed as components of each NatureServe Ecological System. Association data from the NatureServe Explorer API.

State ranks: Conservation status ranks assigned by NatureServe member programs in each state where the ecosystem occurs.