Veronicastrum virginicum

(L.) Farw.

Culver's-root

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G4Apparently SecureGlobal Rank
Identity
Unique IDELEMENT_GLOBAL.2.138942
Element CodePDSCR21010
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVascular Plant
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomPlantae
PhylumAnthophyta
ClassDicotyledoneae
OrderPlantaginales
FamilyPlantaginaceae
GenusVeronicastrum
Other Common Names
Culver's root (EN) Véronique de Virginie (FR)
Concept Reference
Kartesz, J.T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. 2nd edition. 2 vols. Timber Press, Portland, OR.
Taxonomic Comments
Widespread, generally accepted species.
Conservation Status
Rank MethodExpertise without calculation
Review Date2016-08-16
Change Date2000-02-07
Edition Date2000-01-07
Edition AuthorsBarbara S. Dyck
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences81 to >300
Rank Reasons
Species is common, abundant, and widespread in parts of its range in the U.S. In other parts of its range, however, populations are very low. Activities such as habitat conversion to agricultural use, clearing of wooded areas, and drainage/alteration of wet or moist prairies threaten its long term survival, and thus may result in the extirpation of native plants of this species in these areas. In addition, the species is reported to be declining even in states in its core range.
Range Extent Comments
South-eastern Manitoba and south-western Ontario to Maine, south to Florida and Texas (Scoggan 1978-9, Kartesz 1999). Species is recorded in the following provinces and states: Manitoba, Ontario, Alabama, Arkansas, Connecticut, Delaware, Florida, Georgia, Illinois, Indiana, Iowa, Kansas, Kentucky, Louisiana, Maine, Maryland, Massachusetts, Michigan, Minnesota, Mississippi, Missouri, Nebraska, New Jersey, New York, North Carolina, North Dakota, Ohio, Oklahoma, Pennsylvania, South Carolina, South Dakota, Tennessee, Texas, Vermont, Virginia, West Virginia, and Wisconsin.

More detailed range information was available for the following provinces and states.

Manitoba: occurs in the South-Eastern Lake Terrace region (Glacial lake Agassiz Beaches, between the Red River Valley and the Precambrian Shield) from the Manitoba-Minnesota border north to the Kleefeld area (<1% of the area of the province) (E. Punter pers. comm.).

Ontario: native only in southwestern Ontario (Essex, Kent, and Lambton counties) (M. Oldham pers. comm.).

Delaware: occurs in the piedmont and coastal plain (B. McAvoy pers. comm.).

Florida: occurs in the most north-western county in the state (Wunderlin et al. 1995).

Georgia: reported in 2 counties in northern part of the state (USDA, NRCS 1999).

Iowa: infrequent in north-west of state, common elsewhere (J. Peason pers. comm.).

Kansas: largely restricted to the eastern 1/5 of Kansas (eastern 3 tiers of counties) (C. Freeman pers. comm.).

Kentucky: reported in 33 counties scattered throughout the state (USDA, NRCS 1999).

Indiana: occurs in remnants mostly in the northern half of the state but also occurs in the far southern part of the state (Indiana Natural Heritage Data Center).

Massachusetts: occurs in 4 counties in western half of the state (USDA, NRCS 1999).

Maine: apparently introduced to Maine (Haines and Vining 1998), known to occur only in York county (Maine Natural Areas Program).

Michigan: common in southern Lower Michigan (Voss 1985, Voss 1996, Michigan Natural Features Inventory).

Minnesota: occurs in half the state (Minnesota Natural Heritage Program).

Missouri: occurs statewide (Steyermark 1963, M. McHale pers. comm., T. Smith pers. comm.).

North Carolina: recorded in 10 counties in the mountains and piedmont (Radford et al. 1968, J. Amoroso, pers. comm)

Ohio: occurs throughout Ohio, more frequent in the eastern half of the state (A. Cusick pers. comm.).

South Carolina: reported in two counties in the north-west corner and north-central portions of the state (Boyle et al. n.d.).

South Dakota: restricted to southeastern part of state, documented in three counties: Lincoln, Minnehaha, and Moody. Historically it probably occurred in eight counties (D. Ode pers. comm.).

Tennessee: known to occur in 22 counties in Tennessee (APSU Center for Field Biology and University of Tennessee Herbarium 1999).

Virginia: reported in 28 counties mostly in western 2/3 of the state (USDA, NRCS 1999).

Vermont: one current site in the town of Essex, Cittenden County. One historical record last observed in a vacant lot in the City of Rutland. These records may both be garden escapes (R. Popp pers. comm.).

West Virginia: occurs in 18 counties throughout the state (USDA, NRCS 1999).

Wisconsin: occurs in all but the northern-most tier of counties in Wisconsin (K. Westad pers. comm.).

Confirmation was received that the species does not occur in Alberta (Alberta Natural Heritage Information Centre), British Columbia (British Columbia Conservation Data Centre), Quebec (Quebec Service de la Conservation des Especes Menacees), Saskatchewan (Saskatchewan Conservation Data Centre), Alaska (Alaska Natural Heritage Program), Arizona (Arizona Natural Heritage Program), California (California Natural Diversity Database), Montana (Montana Natural Heritage Program), Nevada (Nevada Natural Heritage Program), New Mexico (New Mexico Natural Heritage Program), Utah (Welsh et al. 1993, Utah Natural Heritage Program), Washington (Washington Natural Heritage Program), and Wyoming (Wyoming Natural Diversity Database).
Occurrences Comments
1000's. Manitoba: 12 (E. Punter pers. comm.); Ontario: ~ 15 (M. Oldham pers. comm.); Delaware: 2 (B. McAvoy pers. comm.); Florida: at least 1* (Wunderlin et al. 1995); Georgia: at least 2* (USDA, NRCS 1999); Kansas: 100's (C. Freeman pers. comm.); Kentucky: at least 33* (USDA, NRCS 1999); Missouri: 1000's (T. Smith pers. comm.); Ohio: probably 1000's (A. Cusick pers. comm.); South Dakota: 3+ (D. Ode pers. comm.); Tennessee: at least 22* (APSU Center for Field Biology and University of Tennessee Herbarium 1999); Virginia: at least 28* (USDA, NRCS 1999); Vermont: 1? (R. Popp pers. comm.); Washington: at least 18* (USDA, NRCS 1999); Wisconsin: 1000's? (K. Westad pers. comm.). * signifies a minimum number of populations based on the number of counties for which the species is recorded according to state distribution maps.

It is difficult to distinguish between native and introduced occurrences (W. Moorhead pers. comm.).

Occurences in or near native prairie habitat and away from houses, within its known geographic range, can likely be considered native (B. Ford pers. comm.).

Delaware: there are few populations due to loss of habitat (B. McAvoy pers. comm.).

Louisiana: there are two historic records of this species. It is not known whether these populations have become extirpated (D. Brunet pers. comm.).

North Carolina: this species is on the Natural Heritage Program Watch list because it is rare and poorly known (J. Amoroso pers. comm.).

South Dakota: Ninety percent of its habitat has been destroyed. There may be a few more populations, but there has not been any systematic survey for this species. The number of sites would certainly not total more than 20 (D. Ode pers. comm.).

Vermont: The only extant population was first observed in 1962. In 1982 there were about 20 individuals. In 1991 there were 7 clumps consisting of about 20 stems. In 1999 no plants were observed.

Species is ranked S1 in Manitoba because there are few occurrences known in a small area of the province. Species is being considered for the Endangered Species List in Manitoba (J. Greenall pers. comm.).
Threat Impact Comments
There is both direct and indirect evidence of wild-collection of this species for the plant trade, from Manitoba, Missouri, and Wisconsin.

Manitoba: direct evidence of collecting from the wild observed. Seed or plants must have been taken from the Manitoba Tall Grass Prairie Preserve, since that was the only known population at the time. Two native plant nurseries in Manitoba sell V. virginicum plants which were derived from seed. Probably no rhizomes are collected, since they appear to have dubious medicinal qualities. There is no evidence that this plant is collected from the wild for medicinal purposes. There is no regulation or licensing for wild plant collecting in Manitoba, so no statistics are available (E. Punter pers. comm.)

Missouri: indirect evidence of collecting from the wild. There is a market for the roots but whether it is collected from the wild is unknown. If it is being collected it is not affecting wild populations to any great degree (T. Smith pers. comm.).

Wisconsin: indirect evidence of collecting from the wild (K. Westad pers. comm.).

The plant receives minor but continual usage for medicinal purposes, and it is estimated that 1000-2000 lb/yr were in U.S. trade in the early 1990s (M. McGuffin pers. comm.). The root is used, so harvest is deadly to the plant.

Seed and potted plants of the native form of this species (supposedly) is sold for prairie restoration or native prairie gardening. Plant material is known to have been collected from the wild for this purpose in Manitoba, but the origin of this material in other areas is unknown.

Across its range, Veronicastrum virginicum is reported to be threatened by the following activities:

Conversion of prairie habitat to agricultural use and urban development (A. Cusick pers. comm., B. McAvoy pers. comm., M. Oldham pers. comm.).

Agricultural activities, i.e., pesticide application (E. Punter pers. comm.).

Alteration of hydrology in wet or moist prairie (M. McHale pers. comm.).

Clearing of woodlots (E. Punter pers. comm.).

Road maintenance activities including grading, herbiciding (E. Punter pers. comm.).

Grazing by cattle and white-tail deer (E. Punter pers. comm.).

Removal of shrubs along fence lines and shelterbelts (E. Punter pers. comm.).

Impact of cultivars on native genetic integrity (E. Punter pers. comm.).

Effect of herbicides/insecticides on pollinators (E. Punter pers. comm.).

Low seed production (Lakela 1960, E. Punter pers. comm.).

Loss of open habitat to invasive weeds and pasture grasses (D. Ode, pers.com.), intensive management, and/or reforestation (W. Moorhead pers. comm.).

Encroachment of woody vegetation due to suppression of fires (C. Freeman pers. comm., M. McHale pers. comm.).

Conversion of bogs to wet meadows (J. Amoroso pers. comm.)
Ecology & Habitat

Habitat

Veronicastrum virginicum occurs in a variety of habitats throughout its range. It is found in moist tallgrass prairie and prairie remnants, moist woods, woodland borders, thickets, fields and meadows, stream banks and terraces (Radford et al. 1968, J. Amoroso pers. comm., Maine Natural Areas Program, C. Freeman pers. comm., Pennsylvania Natural Diversity Inventory, B. McAvoy pers. comm., Iowa Department of Natural Resources). It also occurs in secondary habitat on roadsides, road allowances, and railway right-of-ways (A. Cusick pers. comm., Michigan Natural Features Inventory, E. Punter pers. comm., M. Oldham pers. comm., T. Smith pers. comm., K. Westad pers. comm.).

In the north-westernmost part of its range, it occurs in the ecotone between tall grass plant communities and adjoining open to closed deciduous forest, on strongly calcareous, well to imperfectly drained, Dark Grey Chernozemic sandy loam soils (E. Punter pers. comm.). It is found in open oak woodlands in North Dakota (D. Ode pers. comm.), in savanna in Missouri and Wisconsin (M. McHale pers. comm., T. Smith pers. comm., K. Westad pers. comm.), and occasionally in bottomland forest in Missouri (M. McHale, pers.comm.). In North Carolina it is known to occur in bogs, but is primarily found in the mountains (Radford et al. 1968, J. Amoroso pers. comm.). In Wisconsin, it is additionally reported to be found in sand dunes and sedge meadow (K. Westad pers. comm.).

Ecology

Appears to be pollinated by several bee species (E. Punter pers. comm.).
Other Nations (2)
United StatesN4
ProvinceRankNative
NebraskaS1Yes
IowaS4Yes
IndianaSNRYes
New JerseyS3Yes
ArkansasSNRYes
PennsylvaniaS5Yes
IllinoisSNRYes
KentuckyS3Yes
MinnesotaSNRYes
West VirginiaS4Yes
VirginiaS4Yes
South DakotaS1Yes
MaineSNANo
District of ColumbiaS1Yes
GeorgiaS3Yes
TexasSNRYes
North CarolinaS2Yes
North DakotaSHYes
ConnecticutSUYes
MarylandSNRYes
MississippiS1Yes
MissouriSNRYes
OklahomaS1Yes
OhioSNRYes
FloridaSNRYes
MichiganSNRYes
MassachusettsS2Yes
New YorkS2Yes
South CarolinaS1Yes
DelawareS1Yes
WisconsinSNRYes
VermontS1Yes
TennesseeSNRYes
AlabamaS1Yes
KansasS3Yes
LouisianaSHYes
CanadaN2
ProvinceRankNative
Nova ScotiaSNANo
ManitobaS1Yes
OntarioS2Yes
Plant Characteristics
DurationPERENNIAL
Economic Value (Genus)Yes
Roadless Areas (7)
Georgia (1)
AreaForestAcres
Wilson CoveChattahoochee National Forest545
North Carolina (1)
AreaForestAcres
Overflow CreekNantahala National Forest3,379
Virginia (3)
AreaForestAcres
Brush MountainJefferson National Forest6,002
JerkemtightGeorge Washington National Forest16,687
Little AlleghanyGeorge Washington National Forest10,215
West Virginia (2)
AreaForestAcres
Cheat MountainMonongahela National Forest8,191
Cranberry AdditionMonongahela National Forest11,123
References (38)
  1. Amoroso, Jame. Personal communication. Botanist. North Carolina Heritage Program, NC Dept. of Environment, Health, &amp; Natural Resources, Division of Parks And Recreation, Raleigh, NC.
  2. APSU Center for Field Biology and University of Tennessee Herbarium. 1999. October 6-last update. Atlas of Tennessee Vascular Plants. Online. Available: http://www.bio.utk.edu/botany/herbarium/vascular/atlas.html. Accessed 2000-Jan.
  3. Beckstrom-Sternberg, S.M., D.E. Moerman, and J.A. Duke. 1995. Medicinal plants of native America database. Online. Available: http://ars-genome.cornell.edu/cgi-bin/WebAce/webace?db=mpnad b. Accessed 2000, January 7.
  4. Boyle, K., C. Eastman, and T. Mousseau. South Carolina plant atlas. Online. Available: http://cricket.biol.sc.edu/herb. Accessed 2000, January 7.
  5. Brunet, David. Botanist, Louisiana Natural Heritage Program. Personal communication.
  6. Cooperrider, T.S. 1995. The Dicotyledoneae of Ohio. Ohio State University Press, Columbus.
  7. Cusick, A. Botanist, Ohio Natural Heritage Program. Personal communication.
  8. Davidson, Campbell. Director, Morden Research Centre, Agriculture and Agri-Food Canada. Personal communication.
  9. Fernald, M. L. 1950. Gray's manual of botany. 8th edition. Corrected printing (1970). D. Van Nostrand Company, New York. 1632 pp.
  10. Flora of North America Editorial Committee (FNA). 2019. Flora of North America north of Mexico. Vol. 17: Magnoliophyta: Tetrachondraceae to Orobanchaceae. Oxford Univ. Press, New York. xxiv + 737 pp.
  11. Ford, B. Personal communication. Professor and Curator of the University of Manitoba Herbarium, University of Manitoba, Winnipeg, MB.
  12. Freeman, Craig. Personal communication. Botanist, Kansas Natural Features Inventory. Kansas Biological Survey, Lawrence, KS.
  13. Gleason, H.A. 1952. The new Britton and Brown illustrated flora of the northeastern United States and adjacent Canada. 3 volumes. Hafner Press, New York. 1732 pp.
  14. Greenall, Jason. Botanist, Manitoba Department of Natural Resources. Personal communication.
  15. Haines, A. and T.F. Vining. 1998. Flora of Maine, A Manual for Identification of Native and Naturalized Vascular Plants of Maine. V.F.Thomas Co., Bar Harbor, Maine.
  16. Kartesz, J.T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. 2nd edition. 2 vols. Timber Press, Portland, OR.
  17. Kartesz, J.T. 1999. A synonymized checklist and atlas with biological attributes for the vascular flora of the United States, Canada, and Greenland. First edition. In: Kartesz, J.T., and C.A. Meacham. Synthesis of the North American Flora, Version 1.0. North Carolina Botanical Garden, Chapel Hill, N.C.
  18. Kindscher, K. 1992. Medicinal wild plants of the prairie: An ethnobotanical guide. University Press of Kansas, Lawrence.
  19. Lakela, O. 1965. A Flora of Northeastern Minnesota. University of Minnesota Press, Minneapolis, Minnesota. 541 pp.
  20. Looman, J. and K.F. Best. 1979. Budd's flora of the Canadian prairie provinces. Minister of Supply and Services Canada, Ottawa, Ontario.
  21. McAvoy, Bill. Personal communication. Botanist, Delaware Natural Heritage Program.
  22. McGuffin, Michael. Personal communication. American Herbal Products Association, Silver Spring MD
  23. McHale, Maureen. Natural Features Inventory Biologist, The Nature Conservancy/Missouri Department of Conservation. Personal communication.
  24. Moerman, D.E. Native American Ethnobotany: a database of foods, drugs, dyes and fibers of Native American peoples, derived from plants. University of Michigan, Dearborn, MI. Online. Available: http://www.umd.umich.edu/cgi-bin/herb
  25. Moorhead, William. Botanist, Connecticut Natural Diversity Database. Personal communication.
  26. Ode, David. Botanist/Ecologist, South Dakota Natural Heritage Database. Personal communication.
  27. Oldham, Michael. Personal communication. Botanist, Ontario Natural Heritage Information Centre.
  28. Popp, Robert. Heritage Coordinator/Botanist, Vermont Nongame and Natural Heritage. Personal communication.
  29. Punter, C. Elizabeth. Special Projects Botanist, Manitoba Conservation Data Centre. Personal communication.
  30. Radford, A.E., H.E. Ahles, and C.R. Bell. 1968. Manual of the vascular flora of the Carolinas. Univ. North Carolina Press, Chapel Hill, NC. 1183 pp.
  31. Scoggan, H.J. 1978-1979. The flora of Canada: Parts 1-4. National Museums Canada, Ottawa. 1711 pp.
  32. Smith, Tim. Personal communication. Botanist, Missouri Department of Conservation. Missouri Department of Conservation, Jefferson City, MO
  33. Steyermark, J.A. 1963. Flora of Missouri. Iowa State Univ. Press, Ames. 1728 pp.
  34. U.S. Department of Agriculture. 1997. July 16-last update. Western wetland flora: Field office guide to plant species. Online. Available: http://www.npwrc.usgs.gov/resource/othrdata/westflor/westflor.htm. Accessed 2000, January 7.
  35. Voss, E.G. 1985. Michigan flora. Part II. Dicotyledons. Cranbrook Institute of Science and University of Michigan Herbarium. Ann Arbor, Michigan. 1212 pp.
  36. Voss, E.G. 1996. Michigan Flora. Part III. Dicots (Pyrolaceae-Compositae). Cranbrook Institute of Science Bulletin 61 and Univ. Michigan Herbarium. Ann Arbor, Michigan. 622 pp.
  37. Welsh, S.L., N.D. Atwood, S. Goodrich, and L.C. Higgins (eds.) 1993. A Utah flora. 2nd edition. Brigham Young Univ., Provo, Utah. 986 pp.
  38. Westad, Kristin. Wisconsin Natural Heritage Inventory. Personal communication.