Plethodon idahoensis

Slater and Slipp, 1940

Coeur d'Alene Salamander

G4Apparently Secure Found in 8 roadless areas NatureServe Explorer →
G4Apparently SecureGlobal Rank
Least concernIUCN
UnknownThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.100555
Element CodeAAAAD12270
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassAmphibia
OrderCaudata
FamilyPlethodontidae
GenusPlethodon
Other Common Names
Coeur D'Alene Salamander (EN) Salamandre de Coeur d'Alène (FR)
Concept Reference
Frost, D. R. 1985. Amphibian species of the world. A taxonomic and geographical reference. Allen Press, Inc., and The Association of Systematics Collections, Lawrence, Kansas. v + 732 pp.
Taxonomic Comments
Pelletier et al. (2015) found support for two independently evolving lineages within this species based on eight nuclear genes along with mitochondrial data (Crother 2017).

The Coeur d'Alene salamander has been the subject of taxonomic controversy nearly since its initial discovery. Upon first identification, Slater and Slipp (1940) classified it as a new species, Plethodon idahoensis. Lowe (1950) later reclassified it as a subspecies (idahoensis) of a western Washington Plethodon, Van Dyke's salamander (P. vandykei). Some experts cite biochemical evidence that indicates the Coeur d'Alene salamander is a separate species (Highton and Larson 1979, Howard 1993), whereas others hold that the biochemical evidence is weak, and that insufficient morphological variation exists to justify specific designation (Brodie 1970, Nussbaum et al. 1983). Results of a morphometric analysis led Wilson and Larsen (1999) to support recognition of P. vandykei and P. idahoensis as distinct species. Whether considered a species or a subspecies, the Coeur d'Alene salamander represents a unique genetic resource in Idaho, Montana, and British Columbia and should be managed as such (Howard 1993).
Conservation Status
Rank MethodExpertise without calculation
Review Date2016-05-10
Change Date2002-02-13
Edition Date2002-09-20
Edition AuthorsK. Jurist, J. D. Reichel, E. F. Cassirer, C. R. Groves, and G. Hammerson
Threat ImpactUnknown
Range Extent20,000-200,000 square km (about 8000-80,000 square miles)
Number of Occurrences81 - 300
Rank Reasons
Small range in northern Idaho, western Montana, and southeastern British Columbia; habitat loss and fragmentation are the primary threats; recent surveys have greatly expanded the known range in British Columbia; apparently secure.
Range Extent Comments
The Coeur d'Alene Salamander is a remnant of a once diverse plethodontid salamander fauna in the central Rocky Mountains that was likely reduced by climatic changes over the last 10-14 million years (Nussbaum et al. 1983, Tihen and Wake 1981). The species maintains a disjunct distribution at elevations up to 1,524 m (5,000 ft) in northern Idaho, western Montana, and southeastern British Columbia (Wilson et al. 1997; Wilson and Ohanjanian 2002). The North Fork of the Clearwater and the St. Joe drainages in Idaho, and the lower Clark Fork and Kootenai Rivers in Montana, comprise the core of the species distribution (Wilson and Simon 1987, Genter et al. 1988, Groves 1988). The Selway River drainage comprises the southern limit of the known range in Idaho (Wilson 1990) and Copper Creek and the Moyie River drainage the northern limit (Wilson et al. 1989). In Montana, the southern limit of known distribution is Lake Como Falls in the Bitterroot River drainage (Maxell 2002) and the northernmost population is along the South Fork of the Yaak River (Wilson and Simon 1987; Maxell et al. 2003). Coeur d'Alene Salamanders are currently known from about 15 locations in British Columbia, from 95 km north of Revelstoke to a cluster of locations mostly along the southeast corner of Kootenay Lake (Holmberg et al. 1984; Orchard 1990; Charland 1992; Wilson and Ohanjanian 2002).

Although not all localities of occurrence have been identified, the southern, eastern and western edges of the distribution are likely limited by lack of moisture, discontinuous geological formations, and high temperatures (Wilson and Larsen 1998). The species was probably eliminated from most of British Columbia during the last ice age. The northern limit of current distribution probably represents the extent of successful recolonization of suitable habitat (Lynch 1984).
Occurrences Comments
There are 97 to 192 documented U.S. sites: 164 in northern Idaho (69 if all within 2 km of each other are combined) and 28 in Montana (Cassirer et al. 1994). In British Columbia, as of 2001, there were 44 confirmed sites on 42 distinct watercourses (Ohanjanian 1997a, 1997b, 1998, 1999, 2000; Dulisse 1999; Ohanjanian and Beaucher 2000; Ohanjanian et. al 2001).
Threat Impact Comments
Potential threats to the Coeur d'Alene Salamander include road and trail construction, water diversion projects, pollution, timber harvest, exotic species, fire, and illegal collecting. None of these has yet been documented as a cause of population loss or serious mortality for this species.

Habitat fragmentation is a result of road construction, timber harvest, and fire, and it raises additional concerns for Coeur d'Alene Salamanders related to population genetics. Genetic evidence indicates there is little, if any exchange between widely separated (> 100 km) Coeur d'Alene Salamander populations (Howard 1993). However, the extent of dispersal between adjacent sites is unknown. Because Coeur d'Alene Salamanders are typically semi-aquatic, their movements over terrestrial habitats are restricted, particularly in arid forests or areas with little canopy cover. Small populations tend to lose variation by genetic drift more rapidly than larger populations (Lacy 1987). Likewise, they are more susceptible to inbreeding depression, and catastrophic events (e.g., fire, floods) that eliminate or alter habitat and kill individuals. The apparent small size of many Coeur d'Alene Salamander populations, and their apparent isolation, is cause for concern about their continued viability.
Ecology & Habitat

Description

A small, black lungless salamander with a yellowish throat patch, and a yellow, orange, green, or red dorsal stripe. The stripe usually has scalloped edges. Legs are relatively long with short, slightly webbed toes.

Diagnostic Characteristics

Maximum snout-vent length (SVL) (body length without tail) is 62 mm with only 0.5 to 3.0 intercostal folds between adpressed limbs. It has the fewest costal grooves of all western PLETHODON. It also has more vomerine and premaxillary-maxillary teeth and a relatively shorter tail than other western PLETHODON.

Habitat

Primary habitats are seepages and streamside talus; also inhabits talus far from free water (deep talus mixed with moist soil on well-shaded north-facing slopes); in wet weather, occurs also in leaf litter and under bark and logs in coniferous forest. Terrestrial breeder; eggs presumably are laid in underground rock crevices, although no nest sites have been found in the wild.

All plethodontid salamanders respire through their skin; terrestrial species lose water to the environment through evaporation and are therefore restricted to cool, damp environments. Because Coeur d'Alene salamanders may live in the harshest climate of any northwestern PLETHODON (Nussbaum et al. 1983), they are highly dependent on the thermal and hydric stability provided by wet habitats in otherwise inhospitable surroundings. For this reason, Coeur d'Alene and the closely related Van Dyke's salamanders are closely tied to water and are considered among the most aquatic PLETHODON (Brodie and Storm 1971).

Coeur d'Alene salamanders have been found in three major types of habitat: springs or seeps, waterfall spray zones and edges of streams. Seventy-six percent of known locations are classified as seeps, 6% as waterfalls, and 17% as streams. One percent (two sites) are in abandoned mine shafts. However, the relative number of locations in each type is biased by differences in survey efficiency and probably does not reflect the importance of the different habitats. The abundance of seep locations is at least partly due to the relative ease of surveying roadside seeps. Streams and waterfalls are often less accessible, particularly at night. Salamanders are most difficult to find in streamside habitat, where they are usually observed underneath moist rocks on the banks adjacent to the water. Searches of 30 minutes to find a salamander at a stream site during daylight are not uncommon (Groves 1988).

Coeur d'Alene salamander occurrences are generally located in coniferous forest, but are not restricted to a particular overstory species or aspect. Populations have been found in areas with ponderosa pine (PINUS PONDEROSA), Douglas-fir (PSEUDOTSUGA MENZIEZII), western larch (LARIX OCCIDENTALIS), western red cedar (THUJA PLICATA) and western hemlock (TSUGA HETEROPHYLLA) overstories (Groves 1988) at all aspects.

Ninety percent of 99 Idaho occurrences where habitat data have been collected were in areas of greater than 25% canopy cover and only two (both seeps) were in an area with 10% cover or less. Forest cover may be more important near stream sites than seep sites. Average cover at streamside sites (83% + or - 15%) was significantly greater (Mann-Whitney U, P = 0.005) than at seep locations (57% + or - 5%), although only seven stream sites were measured (Cassirer et al. 1994). Minimum canopy cover measured at stream sites was 42%. Terrain at sites was typically steep, with average slopes of 62% (range 10-90%) (Groves 1988, Wilson 1991).

Known populations occur in association with sharply fractured rock formations (used for underground refugia) from 488 m to 1,524 m in elevation. This fractured rock is often found in the Belt Rock formation but can also occur in talus and in other geologic types (Wilson and Simon 1987, Groves and Cassirer 1989). The species is found in conjunction with both persistent and intermittent surface water. Thus, it is possible to locate salamanders at a wet site in the spring, yet be unable to find any animals at the same site later in the summer when the site is dry on the surface.

Reproduction

Coeur d'Alene salamanders mate above ground in late summer and fall (August-October) and, to a lesser extent, in spring (April and May) (Lynch 1984). After a courtship ritual of an hour or more, the male deposits a spermatophore. The sperm cap is picked up by the female with the cloaca (Lynch and Wallace 1987). Females store sperm up to nine months before fertilizing eggs. An average of six eggs is deposited in April or May. The young emerge in mid-September (Lynch 1984).

Neonates apparently grow more slowly than other PLETHODON. Growth probably occurs in spurts associated with wet weather in the spring and fall. Male Coeur d'Alene salamanders reach sexual maturity at 3.5 years of age and females at 4.5 years, but some individuals may delay breeding. Males mate every year, whereas females mate in alternate years (Lynch 1984).
Terrestrial Habitats
Forest - ConiferBare rock/talus/scree
Palustrine Habitats
Riparian
Other Nations (2)
CanadaN3
ProvinceRankNative
British ColumbiaS3Yes
United StatesN3
ProvinceRankNative
MontanaS2Yes
IdahoS3Yes
Threat Assessments
ThreatScopeSeverityTiming
4 - Transportation & service corridorsHigh (continuing)
4.1 - Roads & railroadsHigh (continuing)
5 - Biological resource useHigh (continuing)
5.3 - Logging & wood harvestingHigh (continuing)
7 - Natural system modificationsHigh (continuing)
7.2 - Dams & water management/useHigh (continuing)

Roadless Areas (8)
Idaho (2)
AreaForestAcres
Bighorn - WeitasNez Perce-Clearwater National Forest254,845
North Lochsa SlopeNez Perce-Clearwater National Forest117,662
Montana (6)
AreaForestAcres
HoodooLolo National Forest105,162
Patricks Knob - North CutoffLolo National Forest16,970
Scotchman Peaks (MT)Kootenai National Forest53,909
Sheep Mountain - StatelineLolo National Forest37,844
South Siegel - South CutoffLolo National Forest13,474
Sundance RidgeLolo National Forest7,558
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