Rana cascadae

Slater, 1939

Cascades Frog

G3Vulnerable Found in 132 roadless areas NatureServe Explorer →
G3VulnerableGlobal Rank
Least concernIUCN
HighThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.103017
Element CodeAAABH01060
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicendemic to a single nation
KingdomAnimalia
PhylumCraniata
ClassAmphibia
OrderAnura
FamilyRanidae
GenusRana
Other Common Names
Cascades frog (EN)
Concept Reference
Frost, D. R. 1985. Amphibian species of the world. A taxonomic and geographical reference. Allen Press, Inc., and The Association of Systematics Collections, Lawrence, Kansas. v + 732 pp.
Taxonomic Comments
MtDNA data suggest that R. aurora, R. cascadae, and R. muscosa form a clade within the R. boylii species group (Macey et al. 2001). Crother (2017) states that the disjunct populations should be investigated with respect to call and molecular parameters.
Conservation Status
Rank Method Rank calculation - Biotics v2
Review Date2022-09-06
Change Date2022-09-06
Edition Date2022-09-06
Edition AuthorsHammerson, G., and E. Gaines (2022)
Threat ImpactHigh
Range Extent20,000-200,000 square km (about 8000-80,000 square miles)
Number of Occurrences81 - 300
Rank Reasons
Moderately small range in mountains from Washington to northern California; many extant populations; common in some areas but has declined in other areas, particularly at the southern edge of the range; the species is vulnerable to multiple widespread threats including introduced species, disease, and pollutants.
Range Extent Comments
Range extends from the Cascade and Olympic mountains of northern Washington south to northern California, USA. Populations in the Olympic Mountains of Washington and the Trinity Alps, Mt. Shasta, and Mt. Lassen areas of California are notably disjunct from the primary distribution along the main Cascade axis (Pearl and Adams 2005). Historical locations at low elevations in Washington suggest that the species formerly may have been more broadly distributed (Leonard et al. 1993). Historical elevational range extended from around 400 to 2,500 meters; it is now found above 600 meters (Hallock and McAllister 2009, Pope et al. 2014). Populations in Lassen National Volcanic Monument at the southern end of the range appear to be extirpated (Pope et al. 2014).
Occurrences Comments
This species is widespread throughout its historical habitat in Washington and can be abundant at some sites (Hallock and McAllister 2009). In Oregon, there are more than 100 occurrences, though not all of these are extant, and only 11 of them have a last observed date after 2000 (Pearl et al. 2009, Duarte et al. 2021, ORBIC 2022). Pope et al. (2014) reports 11 populations remain in California, mostly in the Klamath Mountains. It is apparently extirpated from Lassen Volcanic National Park, near the southern end of the species' range (Pope et al. 2014).
Threat Impact Comments
Threats are not fully understood, but their effects are likely interactive and vary by life stage (Alton and Franklin 2017, Cook et al. 2022). Threats are most well-documented at the southern end of the species' distribution in northern California, where declines have been the greatest (Piovia-Scott et al. 2011, Pope et al, 2014, Cole et al. 2016, Alton and Franklin 2017, Evelyn and Sweet 2018, Cook et al. 2022). Introduced predatory fish, habitat loss and degradation (from fire suppression and/or grazing), disease, UV-B radiation, air borne pollutants, and climate change are the greatest threats to this species (Fellers and Drost 1993, Pope et al. 2014, Cole et al. 2016, Evelyn and Sweet 2018, Duarte et al. 2021, Cook et al. 2022).

Livestock grazing can degrade habitat and alter hydrology, reducing available breeding habitat and reducing survival to metamorphosis (Pope et al. 2014, Cole et al, 2016). Grazing is widespread in the range of the Cascades frog, but numbers of livestock have been reduced compared to historical numbers (Pope et al. 2014). Grazing can degrade habitat by increasing sedimentation (Oregon Department of Fish and Wildlife 2016).
Higher elevation sites receive heavy recreational use, altering vegetation and soil structure, increasing sedimentation and modifying mudflats (Pope et al. 2014). Alpine meadow habitats are particularly prone to trampling (Pope et al. 2014).

Fire suppression activities, including drawing water from ponds, application of fire retardant, and construction of fuel breaks may harm populations, but the impact is unclear (Pope et al. 2014, Evelyn and Sweet 2018). Dense forests with high fuel loads as a result of fire suppression can lead to catastrophic wildfire, decreased downslope water inputs, altered peak flows, and increased sedimentation (Pope et al. 2014). Fire suppression on National Forests may have reduced open meadow habitat (Fellers and Drost 1993, Evelyn and Sweet 2018).

Chytridiomycosis, caused by Batrachochytrium dendrobatidis (Bd), is found in Cascades frog populations throughout California and reduces juvenile survival (Piovia-Scott et al. 2011, Pope et al. 2014). The effect of Bd on juvenile survival varies by habitat type, suggesting an interaction between habitat, climate, and disease (Cook et al. 2022).

Egg masses are susceptible to mass die offs from water mold (Saprolegnia ferax) (Blaustein et al 1994, Kiesecker and Blaustein 1997). Nonnative fish are widespread rangewide and prey on Cascades frogs and compete with them for food resources (Pope et al. 2014, Cole et al, 2016, Thurman et al. 2022). Cascades frogs tend not to breed in wet meadows with nonnative trout present (Cole et al. 2016). The effects of predatory fish may also be interactive with other stressors such as climate change and disease (Pope et al. 2014). Removal of introduced fish resulted in increased apparent abundance of Cascades frog adults and larvae in Mount Rainier National Park (Larson et al. 2017).

Pollution is identified as a threat in in Oregon and Washington (Thurman et al. 2022). Air-borne pesticides drifting from California's Central Valley pose a threat to California populations, particularly those in the southern Cascades (Davidson et al. 2002, Pope et al. 2014). However, Davidson et al. (2012) found no difference in pesticide concentrations between these groups, and no correlation with population declines. Increased ultraviolet-B radiation due to ozone depletion is a threat, particularly at higher elevations (Alton and Franklin 2017). This threat interacts with other environmental factors, complicating threat assessment, but contributes to reduced hatching success, larval survival, and activity levels (Kiesecker and Blaustein 1995, Fite et al. 1998, Blaustein et al. 1994, Alton and Franklin 2017).

Climate change will result in altered hydroperiods (earlier drying) in Cascades frog breeding habitat. More severe winters may lead to reduced overwinter survival (Pope et al. 2014, Thurman et al. 2022). This threat is likely greatest in California's southern Cascades, where populations are already small and isolated (Pope et a. 2014). Increased drought associated with climate change will result in altered hydroperiods and reduced habitat for Cascades frogs (Pope et al. 2014, Oregon Department of Fish and Wildlife 2016, Thurman et al. 2022), Increases in drought may affect susceptibility to Bd (Cook et al. 2022). In northern California, shortened hydroperiods were strongly associated with reduced survival to metamorphosis (Cole et al. 2016).
Ecology & Habitat

Description

Most adults are brown to olive-brown and have many distinct black spots on the upper side (black flecks between the spots are scarce or absent), a long yellowish stripe on the upper lip, and a blackish "mask" across the eye and eardrum. Each side of the back has a distinct ridge (dorsolateral fold). The lower abdomen and underside of hind legs are yellow, orange-yellow, or yellowish tan. The groin is usually bright yellow with dark mottling. Lower sides are yellowish or cream. Mature males have a swollen and darkened thumb base and grow to around 60 mm snout-vent length. Females grow larger, to 75 mm. Males produce "chuck-chuck-chuck" or rapid series of clucking calls (several notes per second) during the breeding season; they may call at the water surface or while submerged. Larvae are brown to blackish, with a silvery to brassy underside; they grow to around 5 cm long. Egg masses are rounded clumps of around 300-500 eggs,.deposited singly or often in groups. Primary source: Stebbins (2003).

Diagnostic Characteristics

This species differs from Rana pretiosa in having a less conspicuous light-colored upper jaw stripe, a dark mask, nostrils that are farther apart and lower on the snout, and eyes that are not turned upward as much. It differs from Rana aurora in having distinct black spots on the back, yellowish (rather than red) color on lower abdomen and underside of legs, and generally rougher skin (Stebbins 2003).

Habitat

Cascades frogs inhabit wet mountain meadows, sphagnum bogs, ponds, lakes, and streams, in open or patchy coniferous forests above approximately 600m (Duarte et al. 2021). Breeding sites include wetlands, ponds, lakes, flooded meadows, small alpine ponds and still water edges of small creeks with still water that persists for at least two months after snow melt (Hallock and McAllister 2009). Non-breeding habitat is broader and may include small streams (Pearl and Adams 2005, Pope et al. 2014). Generally they are closely associated with water, but they sometimes may be found in upland habitat during periods of high humidity (Nussbaum et al. 1983). These frogs hibernate in mud at the bottom of ponds and in spring-water saturated ground up to at least 75 meters from a pond (Briggs 1987, Pearl and Adams 2005).

Reproduction

Breeds in spring-summer, March to mid-August, soon after ice and snow melt (Stebbins 1985). In a particular pond, most eggs are laid over a period of just a few days. Each female lays a mass of 300-500 eggs, often in aggregations. Larvae metamorphose into small frogs usually about 2-3 months after the eggs were laid. Individuals first breed probably after their third hibernation (Nussbaum et al. 1983). Typical life span is not more than 5 years.
Terrestrial Habitats
Forest - ConiferGrassland/herbaceous
Palustrine Habitats
TEMPORARY POOLHERBACEOUS WETLANDSCRUB-SHRUB WETLANDBog/fenRiparian
Other Nations (1)
United StatesN3
ProvinceRankNative
WashingtonS3Yes
CaliforniaS3Yes
OregonS3Yes
Threat Assessments
ThreatScopeSeverityTiming
2 - Agriculture & aquacultureLarge (31-70%)UnknownHigh (continuing)
2.3 - Livestock farming & ranchingLarge (31-70%)UnknownHigh (continuing)
2.3.2 - Small-holder grazing, ranching or farmingLarge (31-70%)UnknownHigh (continuing)
6 - Human intrusions & disturbanceRestricted (11-30%)Moderate or 11-30% pop. declineHigh (continuing)
6.1 - Recreational activitiesRestricted (11-30%)Moderate or 11-30% pop. declineHigh (continuing)
7 - Natural system modificationsRestricted (11-30%)Slight or 1-10% pop. declineHigh (continuing)
7.1 - Fire & fire suppressionRestricted (11-30%)Slight or 1-10% pop. declineHigh (continuing)
8 - Invasive & other problematic species, genes & diseasesLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)
8.1 - Invasive non-native/alien species/diseasesLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)
9 - PollutionLarge (31-70%)UnknownHigh (continuing)
9.5 - Air-borne pollutantsLarge (31-70%)UnknownHigh (continuing)
9.5.3 - OzoneLarge (31-70%)UnknownHigh (continuing)
11 - Climate change & severe weatherPervasive (71-100%)Serious - moderateHigh (continuing)
11.2 - DroughtsPervasive (71-100%)Serious - moderateHigh (continuing)

Roadless Areas (132)
California (23)
AreaForestAcres
Bonanza KingShasta-Trinity National Forest16,402
Box CampKlamath National Forest858
Butt Mtn.Lassen National Forest8,217
Castle Crags AShasta-Trinity National Forest113
Castle Crags BShasta-Trinity National Forest1,619
Cub CreekLassen National Forest8,643
Fisher Gulch BShasta-Trinity National Forest4,791
Heart LakeLassen National Forest9,349
JohnsonKlamath National Forest10,652
Lake EleanorShasta-Trinity National Forest397
Little French CShasta-Trinity National Forest11,529
Mill CreekLassen National Forest7,587
Mt. EddyShasta-Trinity National Forest7,232
Orleans Mtn.Klamath National Forest49,090
PortugueseKlamath National Forest18,915
RussianKlamath National Forest21,771
ShacklefordKlamath National Forest1,679
SiskiyouKlamath National Forest54,039
Slate CreekShasta-Trinity National Forest6,636
SnoozerKlamath National Forest23,414
Tom MartinKlamath National Forest9,031
Weaver BallyShasta-Trinity National Forest829
Wild Cattle MtnLassen National Forest4,965
Oregon (46)
AreaForestAcres
Badger CreekMt. Hood National Forest847
BearwallowsDeschutes National Forest7,317
Bend WatershedDeschutes National Forest14,829
Bull Of The WoodsMt. Hood National Forest8,843
Bulldog RockUmpqua National Forest6,056
Bulldog RockWillamette National Forest559
Calf - Copeland CreekUmpqua National Forest15,696
Castle Rock AppendageUmpqua National Forest4,649
Charlton ButteWillamette National Forest3,031
CornpatchWillamette National Forest7,346
Cougar BluffUmpqua National Forest5,574
DoneganUmpqua National Forest5,914
EagleMt. Hood National Forest16,841
Echo MountainWillamette National Forest8,098
French Pete (a)Willamette National Forest1,668
Jackson Creek AppendageUmpqua National Forest4,673
LakeMt. Hood National Forest1,327
LarchMt. Hood National Forest12,961
Limpy RockUmpqua National Forest6,782
Lonesome Meadow AppendageUmpqua National Forest930
Maiden PeakDeschutes National Forest26,432
Maiden PeakWillamette National Forest9,627
Mclennon MountainWillamette National Forest8,085
Menagerie (rooster Rock)Willamette National Forest374
Middle SantiamWillamette National Forest7,316
Mt. JeffersonDeschutes National Forest2,282
Mt. BaileyUmpqua National Forest18,401
Mt. Hood AdditionsMt. Hood National Forest13,061
Mt. ThielsenWinema National Forest1,153
OlallieMt. Hood National Forest7,607
Roaring RiverMt. Hood National Forest27,316
Salmon - HuckleberryMt. Hood National Forest17,570
SherwoodUmpqua National Forest2,449
Skimmerhorn AppendageUmpqua National Forest1,820
Sky Lakes AWinema National Forest3,940
Sky Lakes BWinema National Forest9,615
Thirsty Creek AppendageUmpqua National Forest2,255
Twin LakesMt. Hood National Forest6,055
WaldoDeschutes National Forest4,973
Waldo - FujiWillamette National Forest15,273
Waldo - KochWillamette National Forest245
Waldo - LakeWillamette National Forest2,993
Waldo - Many PrariesWillamette National Forest2,585
Waldo - Salmon CreekWillamette National Forest3,195
West - South BachelorDeschutes National Forest25,994
Wind CreekMt. Hood National Forest5,438
Washington (63)
AreaForestAcres
Alpine Lakes Adj.Wenatchee National Forest57,104
BackboneGifford Pinchot National Forest1,201
Bear CreekGifford Pinchot National Forest7,980
Big Lava BedGifford Pinchot National Forest19,043
Blue LakeGifford Pinchot National Forest11,359
Blue SlideWenatchee National Forest17,505
Boulder RiverMt Baker-Snoqualmie National Forest32,563
BourbonGifford Pinchot National Forest4,512
Canyon CreekWenatchee National Forest7,983
Carlton RidgeGifford Pinchot National Forest2,190
ChambersGifford Pinchot National Forest2,006
ChelanWenatchee National Forest74,650
Coal Creek BluffGifford Pinchot National Forest1,164
CortrightGifford Pinchot National Forest3,462
Dark DivideGifford Pinchot National Forest52,483
Deer CreekGifford Pinchot National Forest882
Devils GulchWenatchee National Forest24,419
Dixon MountainGifford Pinchot National Forest6,339
Eagle RockMt Baker-Snoqualmie National Forest34,064
EntiatWenatchee National Forest72,617
Glacier Peak BMt Baker-Snoqualmie National Forest19,328
Glacier Peak IMt Baker-Snoqualmie National Forest15,175
Glacier Peak JMt Baker-Snoqualmie National Forest26,482
Glacier Peak KMt Baker-Snoqualmie National Forest47,269
Glacier Peak LMt Baker-Snoqualmie National Forest14,084
Goat Rocks AdjWenatchee National Forest6,108
Gotchen CreekGifford Pinchot National Forest7,518
Green MountainOlympic National Forest4,617
Heather LakeWenatchee National Forest10,628
HorseshoeGifford Pinchot National Forest7,579
Jefferson RidgeOlympic National Forest6,512
Jupiter RidgeOlympic National Forest10,148
Liberty BellOkanogan National Forest108,495
LightningOlympic National Forest7,179
Lion RockWenatchee National Forest4,692
ManastashWenatchee National Forest11,155
McdonaldOlympic National Forest501
Moonlight DomeOlympic National Forest4,919
Mt. Baker NorthMt Baker-Snoqualmie National Forest16,873
Mt. BaldyOlympic National Forest3,557
Mt. ZionOlympic National Forest3,544
Nason RidgeWenatchee National Forest19,329
Norse PeakWenatchee National Forest10,169
Packwood LakeGifford Pinchot National Forest1,084
PompeyGifford Pinchot National Forest23,985
QuartzWenatchee National Forest8,550
QuilceneOlympic National Forest18,656
Red LakeGifford Pinchot National Forest2,162
Red MountainGifford Pinchot National Forest2,992
Rock CreekWenatchee National Forest32,239
Rugged RidgeOlympic National Forest4,358
SawtoothOkanogan National Forest122,194
SiouxonGifford Pinchot National Forest12,773
South MidwayGifford Pinchot National Forest2,384
South QuinaultOlympic National Forest11,081
Spencer RidgeGifford Pinchot National Forest5,880
TaneumWenatchee National Forest26,140
TeanawayWenatchee National Forest72,849
Thorp Mtn.Wenatchee National Forest22,717
Twin LakesWenatchee National Forest22,496
Upper SkokomishOlympic National Forest9,311
White PassGifford Pinchot National Forest1,145
WobblyGifford Pinchot National Forest6,335
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