Habitat
Nesting takes place in a variety of habitats including short-grass prairie (Saalfeld et al. 2010, Stocking et al. 2010, Gregory et al. 2011, Coates et al. 2019, Duggar and Duggar 2020), steppe (Stocking et al. 2010), shrub-desert rangeland (Hartman and Oring 2009), pasture (Hartman and Oring 2009, Saalfeld et al. 2010), and agriculture (Saalfeld et al. 2010, Stocking et al. 2010, Duggar and Duggar 2020). Regardless of habitat type, vegetation height is generally below 30 cm (Fellows and Jones 2009, Saalfeld et al. 2010, Stocking et al. 2010, Gregory et al. 2011, Coates et al. 2019, Duggar and Duggar 2020). There is a preference for placing nests near conspicuous objects on the landscape, such as cowpies or rocks, which has been documented in other shorebird species (Coates et al. 2019, Coates et al. 2021). Juveniles use grassland, steppe, pasture, and agricultural habitats with slightly higher average vegetation heights (Fellows and Jones 2009, Hartman and Oring 2009, Stocking et al. 2010).
During migration, habitat use includes dry short-grass prairie, alkali lakes, playa lakes, wet coastal pasture, tidal mudflats, salt marsh, and agricultural fields (Sedgwick 2006, Fellows and Jones 2009, Page et al. 2014, Duggar and Duggar 2020, Coates et al. 2021). In the non-breeding range, a wide variety of coastal and inland habitats are used. Mud flats and salt marshes of tidal estuaries are the primary non-breeding habitat (Sedgwick 2006, Fellows and Jones 2009, Page et al. 2014, Duggar and Duggar 2020, Coates et al. 2021). Agricultural fields have become an increasingly utilized habitat in the non-breeding range in California (Page et al. 2014, eBird 2024), Texas, and northern México (Page et al. 2014, Olalla-Kerstupp et al. 2015b, eBird 2024). Sandy beaches are occasionally used (Duggar and Duggar 2020).
Reproduction
Egg laying begins in April in Washington, by early May in Saskatchewan. Clutch size is 3-5 (usually 4); eggs are laid over 4-7 days. Incubation lasts 28-30 days, by both sexes (Redmond and Jenni 1986). Nestlings are precocial. Young are tended by both parents, brooded at night for several days after hatching. Females usually depart when young are 2-3 weeks old; males tend young until fledging at 41-45 days. Age of first breeding probably is 2-3 years for females, 3-4 years for males. One brood per year. Fledging success is greater for early nesters (Redmond and Jenni 1986). Annual productivity is low. See Allen (1980) for details on nesting and brooding behavior. Often nests in loose colonies. Reported breeding density up to one pair per 24 ha; sometimes only one pair per several hundred ha. Breeding density was about 5-7 males per 100 ha in Idaho (Redmond and Jenni 1986), 1 pair per 6-7 sq km in Saskatchewan, up to 15 territories in 10.4 sq km in Washington (Allen 1980), 0-9 nests per ha in Oregon (Pampush and Anthony 1993), 0.64-2.36 males per sq km in northern Utah (Paton and Dalton 1994). In Oregon, nest success was 0.65-0.69 (Pampush and Anthony 1993). In Utah, 2 of 10 monitored nests were successful (Paton and Dalton 1994).
The likelihood of nest hatching at least one egg has historically been low (Pampus and Anthony 1993, Paton and Dalton 1994). Recent nest success estimates include 14% at 2 sites in Idaho (Coates et al. 2021), 12%-40% at five grassland sites in Iowa and Wyoming (Coates et al. 2019), 29%-31% in Nebraska (Hartman and Oring 2009, Gregory et al. 2011), 23-33% in Oregon and Washington (Stocking et al. 2010). Low nest success is due to nest predation by mammals and birds (Hartman and Oring 2009, Stocking et al. 2010, Gregory et al. 2011, Coates et al. 2019, Coates et al. 2021).