Numenius americanus

Bechstein, 1812

Long-billed Curlew

G4Apparently Secure Found in 55 roadless areas NatureServe Explorer →
G4Apparently SecureGlobal Rank
Least concernIUCN
MediumThreat Impact
Long-billed Curlew (Numenius americanus). Photo by U.S. Fish & Wildlife Service, Public Domain (U.S. Government Work), via ECOS.
U.S. Fish & Wildlife Service, https://www.usa.gov/government-works
Identity
Unique IDELEMENT_GLOBAL.2.100438
Element CodeABNNF07070
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassAves
OrderCharadriiformes
FamilyScolopacidae
GenusNumenius
Other Common Names
Courlis à long bec (FR) long-billed curlew (EN) Zarapito Pico Largo (ES)
Concept Reference
American Ornithologists' Union (AOU). 1998. Check-list of North American birds. Seventh edition. American Ornithologists' Union, Washington, D.C. [as modified by subsequent supplements and corrections published in The Auk]. Also available online: http://www.aou.org/.
Taxonomic Comments
May constitute a superspecies with N. arquata (AOU 1998).
Conservation Status
Rank Method Rank calculation - Biotics v2
Review Date2024-03-12
Change Date2024-03-12
Edition Date2024-03-12
Edition AuthorsGundy, R. L.
Threat ImpactMedium
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences81 - 300
Rank Reasons
This species has wide breeding and non-breeding ranges. It is threatened by habitat loss due to conversion of grassland habitat to agriculture and developed areas, habitat loss due to sea-level rise, and habitat degradation due to invasive species and human modification of the landscape. The population suffered steep historical declines due to overhunting and habitat loss. The population size is around 140,000 individuals and populations are declining by up to 30% throughout the range.
Range Extent Comments
The breeding season (mid-March to mid-July) range is in southwestern Canada and the northwestern United States (Sedgwick 2006, Fellows and Jones 2009, Duggar and Duggar 2020). It extends from southern British Columbia to southwestern Saskatchewan in Canada, southward to northeastern New Mexico, and westward to eastern California, Oregon, and Washington (Sedgwick 2006, Fellows and Jones 2009, Duggar and Duggar 2020). To estimate the breeding range extent, Global Biodiversity Information Facility (GBIF 2024) records were filtered to the years 2004-2024 (i.e., the past 20 years), during the months of May and June (i.e., to exclude as many migratory records as possible), then records that appeared to be outside of the breeding range map in Duggar and Duggar (2020) were excluded. GeoCat (2024) calculated the breeding range extent to be approximately 3.295 million km². The breeding range is the more limited range extent and is the basis for this assessment.

The non-breeding season (mid-July to mid-March) range includes all coastlines of the southern half of the United States, southward through México (including parts of the interior) to the Yucatán Peninsula on the Mexican Gulf Coast and to northern Costa Rica on the Pacific Coast (Sedgwick 2006, Fellows and Jones 2009, Duggar and Duggar 2020). To estimate the non-breeding range extent, GBIF (2024) records were filtered to the years 2004-2024 (i.e., the past 20 years), during the months of November to February (i.e., to exclude as many migratory records as possible), then records that appeared to be outside of the non-breeding range map in Duggar and Duggar (2020) were excluded. GeoCat (2024) calculated the non-breeding range extent to be approximately 9.653 million km².
Occurrences Comments
The exact number of occurrences is unknown. Breeding occurrences likely number in the dozens (Stanley and Skagen 2007, Jones et al. 2008). There are likely several hundred non-breeding occurrences (Sedgwick 2006, Fellows and Jones 2009, Duggar and Duggar 2020, eBird 2024).
Threat Impact Comments
Habitat loss is the greatest threat. The loss of grassland habitat in the breeding range, while happening at a much slower rate than in past centuries, remains a threat (Sedgwick 2006, Fellows and Jones 2009, Duggar and Duggar 2020). Tidal flats, one of the most important non-breeding habitats, have declined 16% worldwide between 1984 and 2016, and the loss of this habitat is projected to continue due to sea-level rise (Murray et al. 2019). Grasslands in the Chihuahuan Desert are declining as they are converted to agriculture (Pool et al. 2014).

Habitat degradation is also occurring. Invasive plants, such as crested wheatgrass (Agropyron cristatum) and diffuse knapweed (Centaurea diffusa), can severely degrade nesting habitat by creating dense, tall stands of vegetation (Sedgwick 2006, Fellows and Jones 2009, Shaffer et al. 2019). Conversely, cheatgrass (Bromus tectorum) has been readily used as a nesting habitat (Sedgwick 2006, Fellows and Jones 2009, Duggar and Duggar 2020). In the absence of native grazing mammals, grassland habitat overgrows and becomes unsuitable without grazing by cattle and sheep or the application of prescribed fire (Sedgwick 2006, Shaffer et al. 2019). Where cattle or sheep graze and where recreational vehicles traverse the landscape, trampling of nests and birds is a localized threat (Sedgwick 2006, Fellows and Jones 2009, Shaffer et al. 2019, Duggar and Duggar 2020). Within agricultural settings, the planting of trees as windbreaks can reduce habitat suitability (Fellows and Jones 2009). Wind farms in the breeding range have the potential to threaten this species by collisions with turbines and the possibility of birds abandoning suitable habitat that is no longer considered attractive with wind turbines in place (Fellows and Jones 2009).

Hunting, a serious historical threat attributed to past declines, is no longer a threat in Canada, the U.S., or México due to legal protections (Sedgwick 2006, Fellows and Jones 2009). Illegal hunting does occur but is generally not considered to be a substantial threat (Sedgwick 2006, Fellows and Jones 2009). However, one study in Iowa found 24 curlew carcasses that had been illegally shot and abandoned by unethical hunters on conservation lands (Katzner et al. 2020).

Pesticides and other contaminants have been detected in this species, but the impacts, if any, are unknown (Fellows and Jones 2009).
Ecology & Habitat

Habitat

Nesting takes place in a variety of habitats including short-grass prairie (Saalfeld et al. 2010, Stocking et al. 2010, Gregory et al. 2011, Coates et al. 2019, Duggar and Duggar 2020), steppe (Stocking et al. 2010), shrub-desert rangeland (Hartman and Oring 2009), pasture (Hartman and Oring 2009, Saalfeld et al. 2010), and agriculture (Saalfeld et al. 2010, Stocking et al. 2010, Duggar and Duggar 2020). Regardless of habitat type, vegetation height is generally below 30 cm (Fellows and Jones 2009, Saalfeld et al. 2010, Stocking et al. 2010, Gregory et al. 2011, Coates et al. 2019, Duggar and Duggar 2020). There is a preference for placing nests near conspicuous objects on the landscape, such as cowpies or rocks, which has been documented in other shorebird species (Coates et al. 2019, Coates et al. 2021). Juveniles use grassland, steppe, pasture, and agricultural habitats with slightly higher average vegetation heights (Fellows and Jones 2009, Hartman and Oring 2009, Stocking et al. 2010).

During migration, habitat use includes dry short-grass prairie, alkali lakes, playa lakes, wet coastal pasture, tidal mudflats, salt marsh, and agricultural fields (Sedgwick 2006, Fellows and Jones 2009, Page et al. 2014, Duggar and Duggar 2020, Coates et al. 2021). In the non-breeding range, a wide variety of coastal and inland habitats are used. Mud flats and salt marshes of tidal estuaries are the primary non-breeding habitat (Sedgwick 2006, Fellows and Jones 2009, Page et al. 2014, Duggar and Duggar 2020, Coates et al. 2021). Agricultural fields have become an increasingly utilized habitat in the non-breeding range in California (Page et al. 2014, eBird 2024), Texas, and northern México (Page et al. 2014, Olalla-Kerstupp et al. 2015b, eBird 2024). Sandy beaches are occasionally used (Duggar and Duggar 2020).

Reproduction

Egg laying begins in April in Washington, by early May in Saskatchewan. Clutch size is 3-5 (usually 4); eggs are laid over 4-7 days. Incubation lasts 28-30 days, by both sexes (Redmond and Jenni 1986). Nestlings are precocial. Young are tended by both parents, brooded at night for several days after hatching. Females usually depart when young are 2-3 weeks old; males tend young until fledging at 41-45 days. Age of first breeding probably is 2-3 years for females, 3-4 years for males. One brood per year. Fledging success is greater for early nesters (Redmond and Jenni 1986). Annual productivity is low. See Allen (1980) for details on nesting and brooding behavior. Often nests in loose colonies. Reported breeding density up to one pair per 24 ha; sometimes only one pair per several hundred ha. Breeding density was about 5-7 males per 100 ha in Idaho (Redmond and Jenni 1986), 1 pair per 6-7 sq km in Saskatchewan, up to 15 territories in 10.4 sq km in Washington (Allen 1980), 0-9 nests per ha in Oregon (Pampush and Anthony 1993), 0.64-2.36 males per sq km in northern Utah (Paton and Dalton 1994). In Oregon, nest success was 0.65-0.69 (Pampush and Anthony 1993). In Utah, 2 of 10 monitored nests were successful (Paton and Dalton 1994).

The likelihood of nest hatching at least one egg has historically been low (Pampus and Anthony 1993, Paton and Dalton 1994). Recent nest success estimates include 14% at 2 sites in Idaho (Coates et al. 2021), 12%-40% at five grassland sites in Iowa and Wyoming (Coates et al. 2019), 29%-31% in Nebraska (Hartman and Oring 2009, Gregory et al. 2011), 23-33% in Oregon and Washington (Stocking et al. 2010). Low nest success is due to nest predation by mammals and birds (Hartman and Oring 2009, Stocking et al. 2010, Gregory et al. 2011, Coates et al. 2019, Coates et al. 2021).
Terrestrial Habitats
Grassland/herbaceous
Palustrine Habitats
HERBACEOUS WETLANDRiparian
Other Nations (2)
CanadaN3B
ProvinceRankNative
AlbertaS3BYes
ManitobaSXBYes
British ColumbiaS4BYes
SaskatchewanS3BYes
United StatesN4
ProvinceRankNative
CaliforniaS2Yes
FloridaS2NYes
UtahS3Yes
WyomingS3BYes
South CarolinaS2NYes
New MexicoS3B,S4MYes
South DakotaS3BYes
MontanaS3BYes
IowaSXBYes
North CarolinaSNAYes
IllinoisSXBYes
Navajo NationS3MYes
WashingtonS2B,S2NYes
KansasS1B,S2NYes
GeorgiaS3Yes
IdahoS2BYes
MinnesotaSXB,SNRMYes
AlabamaS2NYes
LouisianaS3NYes
ColoradoS2BYes
OklahomaS2BYes
NevadaS2BYes
ArizonaS1B,S3NYes
NebraskaS3Yes
North DakotaS2Yes
WisconsinSXBYes
OregonS3BYes
MississippiSNAYes
TexasS3B,S5NYes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)
1.1 - Housing & urban areasLarge (31-70%)Slight or 1-10% pop. declineHigh (continuing)
1.2 - Commercial & industrial areasLarge (31-70%)Slight or 1-10% pop. declineHigh (continuing)
2 - Agriculture & aquacultureLarge - restrictedUnknownHigh (continuing)
2.1 - Annual & perennial non-timber cropsLarge - restrictedUnknownHigh (continuing)
3 - Energy production & miningSmall (1-10%)UnknownHigh - moderate
3.3 - Renewable energySmall (1-10%)UnknownHigh - moderate
5 - Biological resource useSmall (1-10%)Negligible or <1% pop. declineHigh (continuing)
5.1 - Hunting & collecting terrestrial animalsSmall (1-10%)Negligible or <1% pop. declineHigh (continuing)
5.1.3 - Persecution/controlSmall (1-10%)Negligible or <1% pop. declineHigh (continuing)
6 - Human intrusions & disturbanceSmall (1-10%)Negligible or <1% pop. declineHigh (continuing)
6.1 - Recreational activitiesSmall (1-10%)Negligible or <1% pop. declineHigh (continuing)
7 - Natural system modificationsPervasive (71-100%)Slight or 1-10% pop. declineHigh - moderate
7.1 - Fire & fire suppressionPervasive (71-100%)Slight or 1-10% pop. declineHigh - moderate
7.1.2 - Suppression in fire frequency/intensityPervasive (71-100%)Slight or 1-10% pop. declineHigh - moderate
8 - Invasive & other problematic species, genes & diseasesPervasive (71-100%)UnknownHigh (continuing)
8.1 - Invasive non-native/alien species/diseasesPervasive (71-100%)UnknownHigh (continuing)
9 - PollutionPervasive - largeUnknownHigh (continuing)
9.3 - Agricultural & forestry effluentsPervasive - largeUnknownHigh (continuing)
9.3.3 - Herbicides and pesticidesPervasive - largeUnknownHigh (continuing)
11 - Climate change & severe weatherLarge (31-70%)Slight or 1-10% pop. declineHigh (continuing)
11.1 - Habitat shifting & alterationLarge (31-70%)Slight or 1-10% pop. declineHigh (continuing)

Roadless Areas (55)
California (7)
AreaForestAcres
Bear MountainLos Padres National Forest913
Glass MountainInyo National Forest52,867
JuncalLos Padres National Forest12,289
Malduce BuckhornLos Padres National Forest14,177
Santa CruzLos Padres National Forest21,182
Sespe - FrazierAngeles National Forest4,254
TequepisLos Padres National Forest9,080
Idaho (5)
AreaForestAcres
Bear CreekCaribou-Targhee National Forest118,582
Borah PeakSalmon-Challis National Forest130,463
Italian PeakCaribou-Targhee National Forest141,158
Lemhi RangeSalmon-Challis National Forest308,533
PalisadesCaribou-Targhee National Forest122,002
Montana (11)
AreaForestAcres
Bear - Marshall - Scapegoat - SwanLolo National Forest118,485
Bear - Marshall - Scapegoat - SwanLewis and Clark National Forest344,022
East PioneerBeaverhead-Deerlodge National Forest145,082
Freezeout MountainBeaverhead-Deerlodge National Forest97,305
Hyalite - Porcupine - Buffalo Horn Wilderness Study AreaGallatin National Forest143,991
MadisonGallatin National Forest127,859
Middle Mtn. / Tobacco RootsBeaverhead-Deerlodge National Forest96,487
North AbsarokaGallatin National Forest159,075
Snowcrest MountainBeaverhead-Deerlodge National Forest97,649
Tash PeakBeaverhead-Deerlodge National Forest61,312
Ten Lakes #683Kootenai National Forest48,545
Nevada (4)
AreaForestAcres
Bald Mtn.Humboldt-Toiyabe National Forest41,598
Pearl PeakHumboldt-Toiyabe National Forest71,405
Ruby - Secret Pk.Humboldt-Toiyabe National Forest6,749
South SchellHumboldt-Toiyabe National Forest125,614
New Mexico (2)
AreaForestAcres
Candian RiverCibola National Forest7,149
Candian RiverCibola National Forest7,149
North Dakota (8)
AreaForestAcres
Collar / Bennett - CottonwoodDakota Prairie Grasslands19,697
DurlerDakota Prairie Grasslands12,464
DurlerDakota Prairie Grasslands12,464
Long X DivideDakota Prairie Grasslands10,099
SheyenneDakota Prairie Grasslands14,537
SheyenneDakota Prairie Grasslands14,537
VenloDakota Prairie Grasslands5,317
VenloDakota Prairie Grasslands5,317
Oregon (2)
AreaForestAcres
HellholeUmatilla National Forest65,679
Twin MountainWallowa-Whitman National Forest58,533
South Dakota (1)
AreaForestAcres
Indian CreekBuffalo Gap National Grassland24,666
Texas (1)
AreaForestAcres
Big CreekNational Forests in Texas1,447
Utah (4)
AreaForestAcres
North PavantFishlake National Forest53,262
Oak CreekFishlake National Forest54,053
Pine Valley MountainsDixie National Forest57,673
WellsvilleWasatch-Cache National Forest1,717
Wyoming (10)
AreaForestAcres
Grayback RidgeBridger-Teton National Forest295,113
Gros Ventre MountainsBridger-Teton National Forest106,418
Middle ForkShoshone National Forest51,772
Monument RidgeBridger-Teton National Forest17,720
Pacific Creek - Blackrock CreekBridger-Teton National Forest24,658
PalisadesTarghee National Forest1,121
Phillips RidgeBridger-Teton National Forest10,108
West Slope TetonsTarghee National Forest47,448
West Slope WindsBridger-Teton National Forest143,252
Wilderness Study AreaTarghee National Forest51,961
References (72)
  1. Allen, J. N. 1980. The ecology and behavior of the long-billed curlew in southeastern Washington. Wildl. Monogr. 73:1-67.
  2. American Ornithologists' Union (AOU). 1983. Check-list of North American Birds, 6th edition. Allen Press, Inc., Lawrence, Kansas. 877 pp.
  3. American Ornithologists' Union (AOU). 1998. Check-list of North American birds. Seventh edition. American Ornithologists' Union, Washington, D.C. [as modified by subsequent supplements and corrections published in <i>The Auk</i>]. Also available online: http://www.aou.org/.
  4. Andres, B. A., P. A. Smith, R. G. Morrison, C. L. Gratto-Trevor, S. C. Brown, and C. A. Friis. 2012. Population estimates of North American shorebirds, 2012. Wader Study Group Bulletin 119(3):178-194.
  5. Bent, A.C. 1929. Life histories of North American shorebirds (Part II). U.S. Natl. Mus. Bull. 146. Washington, D.C.
  6. BirdLife International. 2004b. Threatened birds of the world 2004. CD ROM. BirdLife International, Cambridge, UK.
  7. BirdLife International. (2013-2014). IUCN Red List for birds. Downloaded from http://www.birdlife.org on various dates in 2013 and 2014. http://www.birdlife.org/
  8. Blus, L. J., C. J. Henny, and A. J. Krynitsky. 1985. Organochlorine-induced mortality and residues in long-billedcurlews from Oregon. Condor 87:563-565.
  9. Carmona, R., N. Arce, V. Ayala-Perez, and G. D. Danemann. 2011. Seasonal abundance of shorebirds at the Guerrero Negro wetland complex, Baja California, Mexico. Wader Study Group Bullet 118(1): 40-48.
  10. Carter, M., C. Hunter, D. Pashley, and D. Petit. 1998. The Watch List. Bird Conservation, Summer 1998:10.
  11. Carter, M., G. Fenwick, C. Hunter, D. Pashley, D. Petit, J. Price, and J. Trapp. 1996. Watchlist 1996: For the future. Field Notes 50(3):238-240.
  12. Coates, S. E., B. W. Wright, and J. D. Carlisle. 2019. Long-billed curlew nest site selection and success in the Intermountain West. The Journal of Wildlife Management 83(5): 1197-1213.
  13. Coates, S., H. Hayes, and J. Carlisle. 2021. IBO long-billed curlew research and community education: 2021 status report. Intermountain Bird Observatory, Boise State University, Idaho. 46 pp.
  14. Cochran, J. F., and S. H. Anderson. 1987. Comparison of habitat attributes at sites of stable and declining long- billed curlew populations. Great Basin Nat. 47:459-466.
  15. Cudworth, N., and A. Orabona. 2012. Evaluating population trends of long-billed curlews in western Wyoming. In Threatened, Endangered, and Nongame Bird and Mammal Investigations, Wyoming Game and Fish Department Nongame Program, Annual Completion Report, edited by: M.B. Grenier, B. Abel, and N. Cudworth.
  16. Dahl, T. E., and S.M. Stedman. 2013. Status and trends of wetlands in the coastal watersheds of the conterminous United States 2004 to 2009. U.S. Department of the Interior, Fish and Wildlife Service and National Oceanic and Atmospheric Administration, National Marine Fisheries Service. 46 pages.
  17. De Smet, K. D. 1992. Status report on the long-billed curlew NUMENIUS AMERICANUS in Canada. Committee on the Status of Endangered Wildlife in Canada (COSEWIC). 28 pp.
  18. Dugger, B. D., and K. M. Dugger. 2020. Long-billed curlew (<i>Numenius americanus</i>), version 1.0. In Birds of the World (A. F. Poole and F. B. Gill, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.lobcur.01
  19. Dugger, Bruce D. and Katie M. Dugger. 2002. Long-billed Curlew (<i>Numenius americanus</i>). The Birds of North America Online (A. Poole, ed.). Cornell Laboratory of Ornithology, Ithaca. Retrieved from the Birds of North America Online: http://bna.birds.cornell.edu.bnaproxy.birds.cornell.edu/bna/species/628/doi:10.2173/bna.628.
  20. eBird. 2024. eBird: An online database of bird distribution and abundance [web application]. eBird, Cornell Lab of Ornithology, Ithaca, New York. Available: http://www.ebird.org.
  21. Ehrlich, P. R., D. S. Dobkin, and D. Wheye. 1992. Birds in Jeopardy: the Imperiled and Extinct Birds of the United States and Canada, Including Hawaii and Puerto Rico. Stanford University Press, Stanford, California. 259 pp.
  22. Fellows, S. D., and S. L. Jones. 2009. Status assessment and conservation action plan for the long-billed curlew (<i>Numenius americanus</i>). U.S. Department of Interior, Fish and Wildlife Service, Biological Technical Publication, FWS/BTP-R60122009, Washington, D.C. 99 pp.
  23. Geospatial Conservation Assessment Tool (GeoCAT). Online. Available: <a href="http://geocat.kew.org/">https://geocat.iucnredlist.org/</a> (Accessed 2024)
  24. Global Biodiversity Information Facility (GBIF). 2024. Global Biodiversity Information Facility (GBIF) data portal. Online. Available: https://www.gbif.org/ (accessed 2024).
  25. Godfrey, W.E. 1966. The birds of Canada. National Museums of Canada. Ottawa. 428 pp.
  26. Gregory, C. J., S. J. Dinsmore, L. A. Powell, and J. G. Jorgensen. 2011. Nest survival of long-billed curlew in Nebraska. Wader Study Group Bulletin 118(2): 109–113.
  27. Harrison, C. 1978. A Field Guide to the Nests, Eggs and Nestlings of North American Birds. Collins, Cleveland, Ohio.
  28. Hartman, C. A., and L. W. Oring. 2009. Reproductive success of long-billed curlews (<i>Numenius americanus</i>) in northeastern Nevada hay fields. Auk 126: 420–430.
  29. Hope, D. D., C. Pekari, M. C. Drever, P. A. Smith, C. Gratto-Trevor, J. Pauquet, Y. Aubry, G. Donaldson, C. Friis, K. Gurney, J. Rausch, A. E. McKellar, and B. Andres. 2019. Shorebirds of conservation concern in Canada - 2019. Wader Study 126(2): 88–100. doi:10.18194/ws.00148
  30. Howell, S. N. G., and S. Webb. 1995. A guide to the birds of Mexico and northern Central America. Oxford University Press, Oxford, UK.
  31. Jehl, J. R., Jr. 1973. Breeding biology and systematic relationships of the stilt sandpiper. Wilson Bulletin 85:115-147.
  32. Jones, S. L., C. S. Nations, S. D. Fellows, and L. L. McDonald. 2008. Breeding abundance and distribution of long-billed curlews (<i>Numenius americanus</i>) in North America. Waterbirds 31(1): 1-14.
  33. Jorgensen, J. G., and S. J. Brenner. 2023. Long-billed curlew (<i>Numenius americanus</i>) surveys in the Nebraska Sandhills 2016 and 2023. Joint Report of the Nongame Bird Program at the Nebraska Game and Parks Commission and Audubon Great Plains, Lincoln, Nebraska, USA. 9 pp.
  34. Katzner, T. E., J. D. Carlisle, S. A. Poessel, E. C. Thomason, B. P. Pauli, D. S. Pilliod, J. R. Belthoff, J. A. Heath, K. J. Parker, K. S. Warner, H. M. Hayes, M. C. Aber, P. A. Ortiz, S. M. Amdor, S. E. Alsup, S. E. Coates, T. A. Miller, and Z. K. Duran. 2020. Illegal killing of nongame wildlife and recreational shooting in conservation areas. Conservation Science and Practice 2020;2:e279. https://doi.org/10.1111/csp2.279
  35. Knopf, F.L. 1996. Mountain Plover (<i>Charadrius montanus</i>). In A. Poole and F. Gill, editors. The Birds of North America, No. 211. The Academy of Natural Sciences, Philadelphia, and The American Ornithologists' Union, Washington, DC. 16 pp.
  36. Knopf, F.L., and J.R. Rupert. 1996. Productivity and movements of mountain plovers breeding in Colorado. Wilson Bulletin 108:28-35.
  37. Morrison, R.I.G. 1993/1994. Shorebird population status and trends in Canada. Bird Trends (3):3-5. Canadian Wildlife Service.
  38. Morrison, R.I.G., McCaffery, B.J., Gill, R.E., Skagen, S.K., Jones, S.K., Page, G.W., Gratto-Trevor, C.L., and Andres, B.A. 2006. Population estimates of North American shorebirds, 2006. Waders Study Group Bulletin 111: 67-85.
  39. Morrison, R. I. G., R. E. Gill, Jr., B. A. Harrington, S. Skagen, G. W. Page, C. L. Gratto-Trevor, and S. M. Haig. 2001. Estimates of shorebird populations in North America. Occasional Paper Number 104, Canadian Wildlife Service, Environment Canada, Ottawa, ON. 64 pages.
  40. Murray, N. J., S. R. Phinn, M. DeWitt, R. Ferrari, R. Johnston, M. B. Lyons, N. Clinton, D. Thau, and R. A. Fuller. 2019. The global distribution and trajectory of tidal flats. Nature 565: 222–225. https://doi.org/10.1038/s41586-018-0805-8
  41. National Audubon Society. 2014. View all Species. Website accessible at http://birds.audubon.org/species. Accessed on various dates in 2014.
  42. Newton, A., J. Icely, S. Cristina, G. M. E. Perillo, R. E. Turner, D. Ashan, S. Cragg, Y. Luo, C. Tu, Y. Li, H. Zhang, R. Ramesh, D. L. Forbes, C. Solidoro, B. Béjaoui, S. Gao, R. Pastres, H. Kelsey, D. Taillie, N. Nhan, A. C. Brito, R. de Lima, and C. Kuenzer. 2020. Anthropogenic, direct pressures on coastal wetlands. Frontiers in Ecology and Evolution 8:144. doi: 10.3389/fevo.2020.00144
  43. Nol, E., and M. S. Blanken. 1999. Semipalmated Plover (<i>Charadrius semipalmatus</i>). No. 444 IN A. Poole and F. Gill, eds. The birds of North America. The Birds of North America, Inc., Philadelphia, PA. 24pp.
  44. Olalla-Kerstupp, A., G. Ruiz-Aymá, J. I. González-Rojas, and A. Guzmán-Velasco. 2015. High fidelity to wintering, stop-over and breeding sites shown by a long-billed curlew <i>Numenius americanus</i> tracked with satellite telemetry on migratory flights across North America. Bird Study 62(4): 556-560. doi: 10.1080/00063657.2015.1073678
  45. Page, G. W., and R. E. Gill, Jr. 1994. Shorebirds in western North America: late 1800s to late 1900s. Studies in Avian Biology 15: 147-160.
  46. Page, G. W., N. Warnock, T. L. Tibbitts, D. Jorgensen, C. A. Hartman, and L. E. Stenzel. 2014. Annual migratory patterns of long-billed curlews in the American West. The Condor 116: 50-61. doi: 10.1650/condor-12-185-r2.1
  47. Pampush, G. J., and R. G. Anthony. 1993. Nest success, habitat utilization and nest-site selection of long-billed curlews in the Columbia Basin, Oregon. Condor 95:957-967.
  48. Parker III, T. A., D. F. Stotz, and J. W. Fitzpatrick. 1996. Ecological and distributional databases for neotropical birds. The University of Chicago Press, Chicago.
  49. Paton, P. W. C., and J. Dalton. 1994. Breeding ecology of long-billed curlews at Great Salt Lake, Utah. Great Basin Nat. 54:79-85.
  50. Pool, D. B., A. O. Panjabi, A. Macias-Duarte, and D. M. Solhjem. 2014. Rapid expansion of croplands in Chihuahua, Mexico threatens declining North American grassland bird species. Biological Conservation 170: 274–281. http://dx.doi.org/10.1016/j.biocon.2013.12.019
  51. Poole, A. F. and F. B. Gill. 1992. The birds of North America. The American Ornithologists' Union, Washington, D.C. and The Academy of Natural Sciences, Philadelphia, PA.
  52. Raffaele, H., J. Wiley, O. Garrido, A. Keith, and J. Raffaele. 1998. A guide to the birds of the West Indies. Princeton University Press, Princeton, NJ. 511 pp.
  53. Redmond, R. L., and D. A. Jenni. 1985. Note on the diet of long-billed curlew chicks in western Idaho. Great Basin Nat. 45:85-86.
  54. Redmond, R. L., and D. A. Jenni. 1986. Population ecology of the long-billed curlew (NUMENIUS AMERICANUS) in western Idaho. Auk 103:755-767.
  55. Ridgely, R. S. 2002. Distribution maps of South American birds. Unpublished.
  56. Ridgely, R. S. and J. A. Gwynne, Jr. 1989. A Guide to the Birds of Panama. 2nd edition. Princeton University Press, Princeton, USA.
  57. Rosenberg, K.V, A.M. Dokter, P.J. Blancher, J.R. Sauer, A.C. Smith, P.A. Smith, J.C. Stanton, A. Panjabi, L. Helft, M. Parr, P.P. Marra. 2019. Decline of the North American avifauna. Science. Published online. 19 Sep 2019.
  58. Rubega, M. A., D. Schamel, and D. M. Tracy. 2000. Red-necked Phalarope (<i>Phalaropus lobatus</i>). No. 538 IN A. Poole and F. Gill, editors, The birds of North America. The Birds of North America, Inc., Philadelphia, PA. 28pp.
  59. Saalfeld, S. T., W. C. Conway, D. A. Haukos, M. Rice, S. L. Jones, and S. D. Fellows. 2010. Multiscale habitat selection by long-billed curlews (<i>Numenius americanus</i>) breeding in the United States. Waterbirds 33(2): 148-161.
  60. Sauer, J. R., J. E. Hines, J. E. Fallon, K. L. Pardieck, D. J. Ziolkowski, Jr., and W. A. Link. 2014. The North American Breeding Bird Survey, Results and Analysis 1966 - 2012. Version 02.19.2014. USGS Patuxent Wildlife Research Center, Laurel, MD. http://www.pwrc.usgs.gov/.
  61. Saunders, E. J. 2001. Population estimate and habitat associations of the long-billed curlew (<i>Numenius americanus</i>) in Alberta. Alberta Species at Risk Report No. 25. Edmonton, Alberta.
  62. Sedgwick, J. A. 2006. Long-billed curlew (<i>Numenius americanus</i>): a technical conservation assessment. USDA Forest Service, Rocky Mountain Region. [Online]. Available: http://www.fs.fed.us/r2/ projects/scp/assessments/longbilledcurlew.pdf [Accessed 11 March 2024].
  63. Senner, N. R. 2006. First record of Long-billed Curlew <i>Numenius americanus</i> in Peru and other observations of Nearctic waders at the Virilla estuary. Cotinga 26:39-42.
  64. Shaffer, J. A., L. D. Igl, D. H. Johnson, M. L. Sondreal, C. M. Goldade, P. A. Rabie, and B. R. Euliss. 2019. The effects of management practices on grassland birds - long-billed curlew (<i>Numenius americanus</i>). Chap. G of Johnson, D.H., Igl, L.D., Shaffer, J.A., and DeLong, J.P., eds., The effects of management practices on grassland birds: U.S. Geological Survey Professional Paper 1842, 12 p., https://doi.org/10.3133/pp1842G.
  65. Sibley, D. A. 2000a. The Sibley guide to birds. Alfred A. Knopf, New York.
  66. Smith, P. A., A. C. Smith, B. Andres, C. M. Francis, B. Harrington, C. Friis, R. I. G. Morrison, J. Paquet, B. Winn, and S. Brown. 2023. Accelerating declines of North America’s shorebirds signal the need for urgent conservation action. Ornithological Applications 125:duad003.
  67. Stanley, T. R., and S. K. Skagen. 2007. Estimating the breeding population of long-billed curlew in the United States. Journal of Wildlife Management 7: 2556-2567.
  68. Stiles, F. G. and A. F. Skutch. 1989. A guide to the birds of Costa Rica. Cornell University Press, Ithaca, New York, USA. 511 pp.
  69. Stocking, J., E. Elliott-Smith, N. Holcomb, and S. M. Haig. 2010. Long-billed curlew breeding success on Mid-Columbia River National Wildlife Refuges, south-central Washington and north-central Oregon, 2007–08: U.S. Geological Survey Open-File Report 2010-1089, 40 pp.
  70. Terres, J. K. 1980. The Audubon Society encyclopedia of North American birds. Alfred A. Knopf, New York.
  71. U.S. Fish and Wildlife Service (USFWS). 1987. Migratory nongame birds of management concern in the United States: the 1987 list. U.S. Fish and Wildlife Service, Office of Migratory Bird Management, Washington, D.C. 63 pp.
  72. Zook, J. L. 2002. Distribution maps of the birds of Nicaragua, Costa Rica, and Panama. Unpublished.