Pipilo erythrophthalmus

(Linnaeus, 1758)

Eastern Towhee

G5Secure Found in 72 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Least concernIUCN
Identity
Unique IDELEMENT_GLOBAL.2.105602
Element CodeABPBX74030
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassAves
OrderPasseriformes
FamilyPasserellidae
GenusPipilo
Other Common Names
eastern towhee (EN) Tohi à flancs roux (FR)
Concept Reference
American Ornithologists' Union (AOU). 1998. Check-list of North American birds. Seventh edition. American Ornithologists' Union, Washington, D.C. [as modified by subsequent supplements and corrections published in The Auk]. Also available online: http://www.aou.org/.
Taxonomic Comments
Formerly regarded as conspecific with P. maculatus (AOU 1998). Citing morphological, behavioral, and mtDNA differences (e.g., Ball and Avise 1992), AOU (1995) split the rufous-sided towhee into two species, P. erythropthalmus (eastern towhee, central and eastern North America) and P. maculatus (spotted towhee, mainly west of the Great Plains). Limited hybridization occurs in a narrow zone in the central Great Plains. See Banks and Browning (1995) for a discussion of nomenclatural issues involving Pipilo.
Conservation Status
Rank MethodExpertise without calculation
Review Date2016-04-09
Change Date1996-12-04
Edition AuthorsPALIS, J.
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Range Extent Comments
BREEDING: southern Manitoba and northeastern North Dakota eastward across the northernmost eastern states and southern Ontario and southwestern Quebec to coastal Maine, south to extreme northeastern Texas (at least formerly) and the Gulf coast states and southern Florida, west to western Iowa, southeastern Nebraska, eastern Colorado, eastern Oklahoma, eastern Arkansas, and eastern Louisiana (AOU 1983, Andrews and Righter 1992, Greenlaw 1996). NON-BREEDING: eastern Kansas, Iowa, southern Great Lakes region, southern New York, and southern New England (rarely farther north) south to southern Texas, Gulf coast, and southern Florida (AOU 1983, Greenlaw 1996).
Threat Impact Comments
Significant declines over last 30 years thought to be primarily the result of forest succession (reduction in early successional habitat). HABITAT CHANGE: Preferred habitat can be lost through vegetation succession or overgrazing. Nearly eliminated from deciduous forest overgrazed by White-tailed Deer (ODOCOILEUS VIRGINIANUS), Elk (CERVUS ELAPHUS), and Mouflon Sheep (OVIS MUSIMON) due to the loss of understory vegetation (Casey and Hein 1983). Abundance was also much lower in unthinned oak woodlands inhabited by numerous White-tailed Deer (13-23/km2) than similar stands inhabited by fewer deer (1-3/km2). Woody understory vegetation was sparse in stands having high deer densities. In thinned stands, however, deer density had no impact on abundance (DeGraaf et al. 1991). Population densities are lower in urbanized areas relative to forested areas due to reduction in suitable habitat (Beissinger and Osborne 1982). Maturation of successional habitats also results in lower population densities (Hagan 1993, Wilcove 1988). NEST PARASITISM: Although forest-dividing corridors can improve habitat conditions for towhees, they also attract Brown-headed Cowbirds (MOLOTHRUS ATER; Rich et al. 1994), which commonly parasitize nests (Friedmann 1963). Frequency of cowbird nest parasitism varies from 3.4% in New York and New Jersey (Greenlaw 1996) to 54.2% in Pennsylvania (Norris 1947). PREDATION: Known nest predators include Pine Snake (PITUOPHIS MELANOLEUCUS), Rat Snake (ELAPHE OBSOLETA), Common Garter Snake (THAMNOPHIS SIRTALIS), and Eastern Chipmunk (TAMIAS STRIATUS). Suspected nest predators include Blue Jay (CYANOCITTA CRISTATA) and weasels (MUSTELA spp.). Known predators of adults include Short-eared Owl (ASIO FLAMMEUS), Sharp-shinned Hawk (ACCIPITER STRIATUS), Cooper's Hawk (ACCIPITER COOPERII), Short-tailed Hawk (BUTEO BRACHYURUS), and Loggerhead Shrike (LANIUS LUDOVICIANUS; Barbour 1951, Greenlaw 1996, Ogden 1974, Storer 1966). OTHER: Migrants are sometimes killed in collisions with towers (Taylor and Anderson 1973).
Ecology & Habitat

Habitat

Inhabits forest and swamp edges, regenerating clearcuts, open-canopied forests (including deciduous, pine [PINUS], pine-hardwood and spruce-fir [PICEA-ABIES]; particularly those with a well-developed understory), reclaimed strip mines, mid-late successional fields, riparian thickets, overgrown fencerows, shrub/small-tree thickets, and other brushy habitats. Nest is typically constructed on the ground, concealed among dense, woody vegetation. Later nests (renests, second broods) are more likely to be built above the ground (usually to 1.5 m, but up to 5.5 m) than earlier nests (Palmer-Ball 1996, Greenlaw 1996). However, regional differences in nest placement occur. Even early nests are more likely to be above ground in the Southeast than in the Northeast (J. Greenlaw, pers. comm.). Habitat selection does not differ appreciably between the breeding and non-breeding seasons (Greenlaw 1996). Significantly more abundant in open-canopied deciduous forest having relatively dense understory growth than in closed-canopied forest with relatively sparse understory vegetation (Annand and Thompson 1997, Chadwick et al. 1986, Childers et al. 1986, Crawford et al. 1981, DeGraaf et al. 1991, Robinson and Robinson 1999, Yahner 1993).

Ecology

DENSITY: In Massachusetts, density of breeding birds varied from an average of 1.78 individuals/ha in closed-canopy forest stands to an average of 7.41 individuals/ha in selectively-harvested stands (Chadwick et al. 1986). In Pennsylvania, the mean number of breeding individuals ranged from 0.17-0.99/ha in six forest types (Yahner 1986). In Arkansas, the number of territorial males ranged from 0.025-0.23/ha. Lowest densities occurred in a burned field and a dry forest, whereas the highest density occurred along a forest edge (James and Neal 1986). In the Pine barrens of southeastern Massachusetts, breeding bird density ranged from about 1.5-3/ha during a three-year period (Morimoto and Wasserman 1991). In New Jersey, population density varied from 0.5 males/ha in mesic oak forest to 0.8 males/ha in xeric pine-oak woodland (Greenlaw 1996). Territory size can vary inversely with experimentally altered food supplies. However, territories appear to contain more than the minimal food stocks necessary for maintenance and reproduction (Wasserman 1983).

INTRA- AND INTERSPECIFIC AGGREGATIONS: Forms loose flocks during the winter, ranging in size from 8-27 (mean = 17) individuals. These wintering groups often form mixed-species foraging flocks with Dark-eyed Juncos (JUNCO HYEMALIS), White-throated Sparrows (ZONOTRICHIA ALBICOLLIS), Field Sparrows (SPIZELLA PUSILLA), Song Sparrows (MELOSPIZA MELODIA), Northern Cardinals (CARDINALIS CARDINALIS), Tufted Titmice (BAEOLOPHUS BICOLOR), Winter Wrens (TROGLODYTES TROGLODYTES) and Carolina Chickadees (POECILE CAROLINENSIS; Barbour 1941).

SITE FIDELITY: Exhibits breeding site fidelity. In New Jersey, one male returned to the same 24-hectare section of oak forest in 5 of 6 years (Greenlaw 1996). In another New Jersey study, the return rate was 20% the first year after banding and 43% thereafter (Leck et al. 1988).

POPULATION PARAMETERS: Sex ratio data are limited; however, sex ratios determined to date are male-biased (at least temporarily) or no different from parity (Greenlaw 1996). Estimated annual probability of survival among adults is 0.58 (Savidge and Davis 1974 cited in Greenlaw 1996). Oldest known individual was at least 12 years, 3 months old (Terres 1991).

PARASITES: Host to at least 14 species of ectoparasites and six internal parasites (Greenlaw 1996).

Reproduction

Both sexes become sexually mature in second year of life (Greenlaw 1996).

PHENOLOGY: Breeding season varies with latitude. In the southeastern U.S. (Florida and Georgia) nest building and oviposition begin in late March-early April, whereas in New England these activities are delayed until mid-May (Greenlaw 1996, Stevenson and Anderson 1994). In the Midwest, breeding activities begin in mid- to late April (Mumford and Keller 1984).

OVIPOSITION/INCUBATION: Dates of oviposition can vary among habitats at the same latitude. In New Jersey, for example, oviposition began earlier in mesic oak forest than in xeric pine barrens. This difference was attributed to earlier foliage development and the concomitant earlier appearance of additional invertebrates in the oak forest (Greenlaw 1978). Egg-laying generally terminates by mid- to late July, but can extend into mid to late August. Clutch size is 2-6 (usually 3-4 eggs), and incubation begins with the laying of the penultimate or ultimate egg (Greenlaw 1996). Clutch size can vary at the same latitude, with larger clutches being produced in habitats with greater food supplies (Greenlaw 1978). Incubation, by the female only, lasts 12-13 days. Both sexes feed the nestlings and fledglings.

FLEDGING: Young leave the nest when 10-11 days old. Produce 1-2 broods per season; will also renest after nest failure (Greenlaw 1996).

NEST SUCCESS: At two sites in New Jersey, egg success (number of eggs that produced fledged young) varied from 28.3-35.6%. Nest predation rates ranged from 31-73% at these two sites (Greenlaw 1996). In West Virginia, nest success (nests that fledged at least one young) was 36.5% (Bell and Whitmore 1997).
Terrestrial Habitats
Woodland - HardwoodWoodland - ConiferWoodland - MixedShrubland/chaparralOld field
Palustrine Habitats
Riparian
Other Nations (2)
CanadaN4B,N3N
ProvinceRankNative
OntarioS4B,S3NYes
QuebecS3BYes
SaskatchewanS3BYes
ManitobaS3BYes
United StatesN5
ProvinceRankNative
MissouriSNRB,SNRNYes
District of ColumbiaS4B,S4NYes
PennsylvaniaS5B,S4N,S4MYes
OklahomaS4Yes
DelawareS5BYes
South DakotaS3BYes
ArkansasS3Yes
MarylandS5B,S4NYes
South CarolinaS5Yes
MassachusettsS4BYes
IndianaS4BYes
IowaS4B,S4NYes
ConnecticutS5BYes
MinnesotaSNRBYes
KentuckyS5B,S5NYes
FloridaS4Yes
MichiganS5Yes
GeorgiaS5Yes
IllinoisS5Yes
MississippiS5B,S5NYes
KansasS3B,S2NYes
New YorkS5BYes
TexasS2B,S5NYes
North CarolinaS5B,S5NYes
AlabamaS5Yes
LouisianaS5Yes
VirginiaS5Yes
West VirginiaS5B,S5NYes
VermontS4BYes
TennesseeS5Yes
Rhode IslandS5BYes
OhioS5Yes
WisconsinS3BYes
New JerseyS4B,S4NYes
MaineS4BYes
New HampshireS4BYes
North DakotaSNRBYes
NebraskaS4Yes
Roadless Areas (72)
Florida (2)
AreaForestAcres
Alexander Springs CreekOcala National Forest2,954
Clear LakeApalachicola National Forest5,592
Georgia (2)
AreaForestAcres
Kelly RidgeChattahoochee National Forest8,325
Lance CreekChattahoochee National Forest9,025
Indiana (1)
AreaForestAcres
Mogan RidgeHoosier National Forest8,435
Kentucky (1)
AreaForestAcres
WolfpenDaniel Boone National Forest2,835
Michigan (1)
AreaForestAcres
Bear SwampHuron-Manistee National Forest3,915
North Carolina (17)
AreaForestAcres
Bald MountainPisgah National Forest11,085
Balsam ConePisgah National Forest10,591
BearwallowPisgah National Forest4,113
Craggy MountainPisgah National Forest2,657
Dobson KnobPisgah National Forest6,111
Graveyard Ridge (addition)Pisgah National Forest1,958
Jarrett CreekPisgah National Forest7,485
Laurel MountainPisgah National Forest5,683
Linville Gorge AdditionPisgah National Forest2,809
Lost CovePisgah National Forest5,944
Middle Prong AdditionPisgah National Forest1,852
Sam Knob (addition)Pisgah National Forest2,576
SnowbirdNantahala National Forest8,489
South Mills RiverPisgah National Forest8,588
Tusquitee BaldNantahala National Forest13,670
Wesser BaldNantahala National Forest4,061
Woods MountainPisgah National Forest9,602
Pennsylvania (3)
AreaForestAcres
Allegheny FrontAllegheny National Forest7,430
Minister ValleyAllegheny National Forest1,417
Tracy RidgeAllegheny National Forest9,034
Tennessee (7)
AreaForestAcres
Bald MountainCherokee National Forest11,743
Bald River Gorge AdditionCherokee National Forest1,728
Beaver Dam CreekCherokee National Forest5,070
Brushy RidgeCherokee National Forest7,469
Flint Mill GapCherokee National Forest9,494
Stone MountainCherokee National Forest5,367
Sycamore CreekCherokee National Forest6,984
Virginia (25)
AreaForestAcres
Adams PeakGeorge Washington National Forest7,135
Bear CreekJefferson National Forest18,274
Beards MountainGeorge Washington National Forest7,505
Beaver Dam CreekJefferson National Forest1,135
Broad RunJefferson National Forest10,971
Brush MountainJefferson National Forest6,002
Elliott KnobGeorge Washington National Forest9,380
Gum RunGeorge Washington National Forest12,620
Hunting Camp Little Wolf CreekJefferson National Forest8,953
JerkemtightGeorge Washington National Forest16,687
Kelley MountainGeorge Washington National Forest7,590
Little Wilson Creek Addition BJefferson National Forest1,725
Mill MountainGeorge Washington National Forest10,840
Mt. PleasantGeorge Washington National Forest8,933
North MountainJefferson National Forest8,377
Northern MassanuttenGeorge Washington National Forest9,444
Oak KnobGeorge Washington National Forest10,882
Peters Mountain Addition BJefferson National Forest2,909
Price MountainJefferson National Forest9,119
Raccoon BranchJefferson National Forest4,388
Saint Marys AdditionGeorge Washington National Forest1,454
Seng MountainJefferson National Forest6,428
Shawvers Run AdditionJefferson National Forest1,927
SkidmoreGeorge Washington National Forest5,641
Southern MassanuttenGeorge Washington National Forest11,985
West Virginia (13)
AreaForestAcres
Canaan LoopMonongahela National Forest7,867
Cranberry AdditionMonongahela National Forest11,123
Cranberry Glades Botanical AreaMonongahela National Forest785
Dolly Sods Roaring PlainMonongahela National Forest13,392
Dry River (WV)George Washington National Forest7,331
Falls Of Hills CreekMonongahela National Forest6,925
Glady ForkMonongahela National Forest3,239
Little MountainMonongahela National Forest8,172
Mcgowan MountainMonongahela National Forest10,504
Middle MountainMonongahela National Forest19,020
North Mountain HopevilleMonongahela National Forest6,525
Seneca CreekMonongahela National Forest22,287
Tea Creek MountainMonongahela National Forest8,295
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