Myotis lucifugus

(Le Conte, 1831)

Little Brown Myotis

G3Vulnerable (G3G4) Found in 16 roadless areas NatureServe Explorer →
G3VulnerableGlobal Rank
EndangeredIUCN
Very high - highThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.100473
Element CodeAMACC01010
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNEndangered
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassMammalia
OrderChiroptera
FamilyVespertilionidae
GenusMyotis
Other Common Names
Little Brown Bat (EN) little brown bat (EN) Petite chauve-souris brune (FR) Un Murciélago (ES) Vespertilion brun (FR)
Concept Reference
Wilson, D. E., and D. M. Reeder (editors). 2005. Mammal species of the world: a taxonomic and geographic reference. Third edition. The Johns Hopkins University Press, Baltimore. Two volumes. 2,142 pp. [As modified by ASM the Mammal Diversity Database (MDD) at https://www.mammaldiversity.org/index.html]
Taxonomic Comments
The five previously recognized subspecies of Myotis lucifugus (alascensis, carissima, lucifugus s.s., pernox, relictus) have been recognized as distinct species by Morales and Carstens (2018). NatureServe will continue to recognize the broader concept of M. lucifugus following the American Society of Mammalogists (ASM 2025) and Bats of the World (Simmons and Cirranello 2025). This follows Francis et al. (2022) who recommend that M. l. alascensis, M. l. carissima, M. l. pernox, and M. l. relictus should continue to be recognized as subspecies of Myotis lucifugus, at least until such time as more conclusive information is provided supporting their elevation to species status.

Myotis occultus was formerly included in M. lucifugus (Jones et al. 1992; Koopman, in Wilson and Reeder 1993), but is now regarded as a distinct species.
Conservation Status
Rank Method Rank calculation - Biotics v2
Review Date2024-06-28
Change Date2021-12-02
Edition Date2024-06-28
Edition AuthorsHunting, K.
Threat ImpactVery high - high
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences> 300
Rank Reasons
This species remains widespread in North America although abundance has decreased in recent years especially in the northeast and some parts of the Midwest where the effects of White-nose Syndrome are currently the most severe. There is some evidence suggesting short-term adaptation to the effects of this disease (Gignoux-Wolfsohn et al. 2021) but more work is needed to determine overall population response. Colony mortality from extreme temperatures and the effects of drought on insect food supply at colony sites are a significant impact in many areas and are likely to become more prevalent as the effects of climate change intensify.
Range Extent Comments
This species occurs in forested regions throughout the northern United States south into parts of central and southern California, Arizona, and New Mexico and as far south as Georgia in the southeastern U.S.; north through Canada, and most of southern and central Alaska (Shively and Barbosa 2017, Tessler et al 2014). It is reported by state wildlife programs in the U.S. to occur in 37 states where it is generally rated as a high priority Species of Greatest Conservation Need in the Midwest and East and lower priority elsewhere. This species is listed as endangered in Maine and Massachusetts (USGS 2024). The North American range of this species was estimated at about 9,763,500 km2 using a concave hull analysis with recent observation locations as inputs.
Occurrences Comments
The number of occurrences of this is unknown but it is represented by a large number of occurrences spread over a vast geographic range.
Threat Impact Comments
The single most significant current threat to this species is mortality from behavioral changes induced by infection from the fungus Pseudogymnoascus destructanans commonly known as White-nose Syndrome. Mortality as high as 90% of colony members in hibernacula have been reported from the eastern U.S. (Cheng et al (2021) and eastern Canada (COSEWIC 2013). Other threats to this species include mortality from wind turbine strikes (direct impact and barotrauma) (O’Shea et al 2016, Choi et al 2022, COSEWIC 2013) although mortality of this species among Canadian bat species appears to be low from wind turbine strikes; intentional (addressing fear of disease spread) and unintentional (demolition of older structures) colony eradication; loss and degradation of foraging habitat that provide an adequate supply of insects; and disturbance of hibernacula although many known sites are now protected. Finally, drought and extreme heat events exacerbated in time and space by climate change are an increasing threat to this and other bat species (Hayes and Adams 2017). Evidence suggests mass lethal hyperthermia events at hibernacula are increasing (see O’Shea et al 2016) and substantial decreases in reproduction during droughts in some species have been empirically demonstrated (Adams 2010). There is some evidence that this species may be rapidly adapting to infection from this fungal pathogen through detected changes in genes associated with the immune system in populations of this species recovering from infection events (Gignoux-Wolfsohn et al 2021).
Ecology & Habitat

Description

Cinnamon-buff to dark brown above, buffy to pale gray below; hairs on back have long glossy tips; ear when laid forward reaches approximately the nostril; tragus about half as high as ear; calcar without keel; length of head and body 41-54 mm, ear 11.0-15.5 mm, forearm 33-41 mm; braincase rises gradually from rostrum; greatest length of skull 14-16 mm; length of upper toothrow 5.0-6.6 mm (Hall 1981).

Diagnostic Characteristics

Differs from M. sodalis in unkeeled calcar. Differs from M. austroriparius in smaller size, glossy rather than dull pelage, and usual absence of a sagittal crest. Differs from M. grisescens in banded dorsal hairs banded (vs. unicolored) and wing attached to the foot at the base of the toe rather than at the ankle. Differs from M. velifer in smaller size, glossy rather than dull pelage, and lack of sagittal crest. Differs from M. keenii and M. septentrionalis in shorter ears that do not extend beyond the nose when laid forward. Differs from M. volans in smaller size, glossy rather than dull pelage, and unkeeled calcar. Differs from M. yumanensis in larger size, larger skull (greatest length usually more than 14 mm rather than usually less than 14 mm), and usually glossy pelage rather than dull pelage. Differs from M. thysanodes in absence of a conspicuous fringe of hairs along the edge of the interfemoral membrane. Differs from M. californicus in larger size, unkeeled calcar, and skull rising gradually from rostrum. Differs from M. leibii in larger size and unkeeled calcar. (Hall 1981).

Habitat

These bats use a wide range of habitats and often use human-made structures for resting and maternity sites; they also use caves and hollow trees. Foraging habitat requirements are generalized; foraging occurs over water, along the margins of lakes and streams, or in woodlands near water. Winter hibernation sites (caves, tunnels, abandoned mines, and similar sites) generally have a relatively stable temperature of about 2-12 C (see Kunz and Reichard 2010). Maternity colonies commonly are in warm sites in buildings (e.g., attics) and other structures; also infrequently in hollow trees. Microclimate conditions suitable for raising young are relatively narrow, and availability of suitable maternity sites may limit the species' abundance and distribution.

Ecology

Winter concentrations may include tens of thousands of individuals. Survival rate is low during the first winter, higher in subsequent years.

Reproduction

Mating occurs usually in September-October. Ovulation and fertilization are delayed until spring. Gestation lasts 50-60 days. Reproductive females annually give birth to 1 litter of 1 young, late spring-early summer. Females produce first young usually in their first (Indiana, New Mexico) or second year (British Columbia) (Herd and Fenton 1983). In British Columbia, individuals may delay or forego reproduction in wet years (Grindal et al. 1992). Survival for a decade may be fairly common; a few live as long as 20-30 years; females may be reproductive to an age of at least 12 years (Hall et al. 1957, Keen and Hitchcock 1980). Most summer colonies range from 50 to 2,500 individuals (average 400) (Mumford and Cope 1964).
Terrestrial Habitats
Forest - HardwoodForest - MixedWoodland - HardwoodWoodland - MixedShrubland/chaparralGrassland/herbaceousOld fieldSuburban/orchardUrban/edificarianAerial
Palustrine Habitats
HERBACEOUS WETLANDFORESTED WETLANDBog/fenRiparianAerial
Other Nations (2)
United StatesN3
ProvinceRankNative
MississippiSHYes
FloridaSNRYes
UtahS3Yes
MontanaS2Yes
MassachusettsS2Yes
District of ColumbiaS4Yes
OklahomaS2Yes
PennsylvaniaS1Yes
KansasS3Yes
IdahoS3Yes
WyomingS5Yes
MichiganS1Yes
NevadaS2Yes
TennesseeS3Yes
ColoradoS4Yes
IndianaS1Yes
MissouriS2Yes
CaliforniaSNRYes
WisconsinS3Yes
Rhode IslandS5Yes
WashingtonS3Yes
MinnesotaS3Yes
OregonS3Yes
KentuckyS2Yes
MaineS5Yes
DelawareS1Yes
ArkansasS1Yes
GeorgiaS1Yes
VirginiaS2Yes
AlabamaS3Yes
ConnecticutS1Yes
VermontS1Yes
New YorkS1Yes
IowaS4Yes
South DakotaS5Yes
OhioS1Yes
MarylandS1Yes
IllinoisS5Yes
South CarolinaS2Yes
NebraskaSNRYes
New JerseyS1Yes
New HampshireS1Yes
West VirginiaS1Yes
AlaskaS3Yes
North CarolinaS2Yes
North DakotaSNRYes
CanadaN3B,NNRN,NNRM
ProvinceRankNative
British ColumbiaS3Yes
Prince Edward IslandS1Yes
LabradorS2Yes
ManitobaS2N,S5BYes
Island of NewfoundlandS2Yes
AlbertaS3B,SNRN,SNRMYes
Northwest TerritoriesS3Yes
Nova ScotiaS1Yes
SaskatchewanS4B,S4NYes
QuebecS1Yes
New BrunswickS1Yes
Yukon TerritoryS3BYes
OntarioS3Yes
Threat Assessments
ThreatScopeSeverityTiming
3 - Energy production & miningRestricted (11-30%)Moderate or 11-30% pop. declineHigh (continuing)
3.3 - Renewable energyRestricted (11-30%)Moderate or 11-30% pop. declineHigh (continuing)
5 - Biological resource useLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)
5.1 - Hunting & collecting terrestrial animalsLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)
5.1.3 - Persecution/controlLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)
6 - Human intrusions & disturbanceRestricted - smallModerate - slightHigh (continuing)
8 - Invasive & other problematic species, genes & diseasesPervasive (71-100%)Extreme - moderateHigh (continuing)
8.1 - Invasive non-native/alien species/diseasesPervasive (71-100%)Extreme - moderateHigh (continuing)
9 - PollutionLarge - smallUnknownHigh (continuing)
11 - Climate change & severe weatherPervasive (71-100%)Moderate or 11-30% pop. declineHigh (continuing)
11.2 - DroughtsLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)
11.3 - Temperature extremesLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)

Roadless Areas (16)
Alaska (7)
AreaForestAcres
College FiordChugach National Forest1,130,818
Freshwater BayTongass National Forest44,933
Game CreekTongass National Forest54,469
Kenai LakeChugach National Forest213,172
North RevillaTongass National Forest215,430
Upper SitukTongass National Forest16,789
Yakutat ForelandsTongass National Forest323,648
California (2)
AreaForestAcres
Hoover - NorthHumboldt-Toiyabe National Forest1,574
Iceberg - Mill CreekHumboldt-Toiyabe National Forest26,988
Idaho (1)
AreaForestAcres
NeedlesPayette National Forest131,279
Montana (2)
AreaForestAcres
Big Snowy Mountains WsaLewis and Clark National Forest88,003
Freezeout MountainBeaverhead-Deerlodge National Forest97,305
New Hampshire (1)
AreaForestAcres
Great Gulf Ext.White Mountain National Forest15,110
Oregon (1)
AreaForestAcres
HomesteadWallowa-Whitman National Forest5,817
Pennsylvania (1)
AreaForestAcres
Tracy RidgeAllegheny National Forest9,034
Utah (1)
AreaForestAcres
418025Uinta National Forest32,698
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