Mylopharodon conocephalus

(Baird and Girard, 1854)

Hardhead

G3Vulnerable Found in 7 roadless areas NatureServe Explorer →
G3VulnerableGlobal Rank
Least concernIUCN
Identity
Unique IDELEMENT_GLOBAL.2.105531
Element CodeAFCJB25010
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicendemic to a single state or province
KingdomAnimalia
PhylumCraniata
ClassActinopterygii
OrderCypriniformes
FamilyLeuciscidae
GenusMylopharodon
Concept Reference
Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea, and W.B. Scott. 1991. Common and scientific names of fishes from the United States and Canada. American Fisheries Society, Special Publication 20. 183 pp.
Taxonomic Comments
See Moyle et al. (1989) for nomenclatural history. Perhaps should be included in the genus Ptychocheilus (Lee et al. 1980), but a morphometric phylogenetic analysis of Mylopharodon and Ptychocheilus by Mayden et al. (1991) concluded that the best arrangement is to retain Mylopharodon as a monotypic genus distinct from Ptychocheilus.
Conservation Status
Rank MethodExpertise without calculation
Review Date2014-09-18
Change Date2014-09-18
Edition Date2012-05-03
Edition AuthorsHammerson, G., T. Hopkins, T., and P. Moyle
Range Extent20,000-200,000 square km (about 8000-80,000 square miles)
Number of Occurrences21 - 80
Rank Reasons
Widely distributed in California but declining and vulnerable to local extirpation from habitat alteration (stream flow quantity and quality changes) and introduced fishes.
Range Extent Comments
Range includes much of the Sacramento-San Joaquin drainage of California (widely distributed in foothill streams), from the Kern River, Kern County, in the south to the Pit River drainage in Modoc County in the north (where widely distributed south of the Goose Lake drainage, and including spotty occurrences in the main Pit River and associated hydroelectric reservoirs); also the Russian River drainage (where uncommon) (Moyle et al. 1989, Moyle 2002). This species occurs in scattered tributaries of the San Joaquin River but not in the valley reaches of the river (Moyle et al. 1989, Moyle 2002). The species is still fairly common in the mainstem Sacramento River, in the lower reaches of the American and Feather rivers, in some smaller tributary streams (e.g., Deer, Pine, and Clear creeks), and in some river reaches above foothill reservoirs (Moyle 2002). The species is absent from the Clear Lake basin and from San Francisco Bay streams except the Napa River (where now extremely rare) (Moyle 2002). Elevational range is 10-1,450 meters. The species is still widespread (though less common, especially in the southern part of the range).
Occurrences Comments
This species is represented by a fairly large number of occurrences (subpopulations) and locations.
Threat Impact Comments
Dams and diversions have eliminated habitat and left many populations isolated and vulnerable to local extinction due to unsuitable stream temperatures and flows (Moyle 2002). Centrarchid fishes (bass, sunfish) threaten populations in foothill streams. Reservoir populations should not be regarded as "safe" populations since these also are vulnerable to large declines caused by increased populations of introduced smallmouth bass and other centrarchid basses (Moyle et al. 1989, Moyle 2002). Population crashes in Pardee, Millerton, Berryessa, Don Pedro, and Folsom reservoirs are thought to have resulted from centrarchid bass predation (see Moyle 2002).
Ecology & Habitat

Habitat

Habitat includes deep, rock- and sand-bottomed pools of small to large rivers (Page and Burr 2011). This species is a good indicator of relatively undisturbed conditions. Usually it occurs in the less disturbed sections of larger low and mid-elevation streams (small to large rivers). It appears to prefer clear, deep pools with sand-gravel-boulder substrates and slow water. It is permanently established in some mid-elevation reservoirs but is usually impermanent in reservoirs (populations may become large, then crash). In streams, it tends to be in the lower half of the water column; it may be found near the surface in reservoirs and in large pools of rivers downstream from reservoirs. This species is always associated with Sacramento pikeminnow and usually with Sacramento sucker. Generally it does not occur in waters dominated by introduced species, especially centrarchids (Moyle et al. 1989). Spawning occurs probably in gravel riffles of smaller tributary streams.

Ecology

Usually found in association with Sacramento squawfish.

Reproduction

Spawning occurs presumably in the spring, perhaps by May-June in Valley streams, extending into August in foothill streams (Moyle et al. 1989). Mature after 2nd year (Moyle 1976).
Other Nations (1)
United StatesN3
ProvinceRankNative
CaliforniaS3Yes
Roadless Areas (7)
California (7)
AreaForestAcres
Bucks LakePlumas National Forest680
Chips CreekLassen National Forest29,089
Chips CreekPlumas National Forest12,940
Devil GulchSierra National Forest30,490
Greenhorn CreekSequoia National Forest28,226
Mill CreekSequoia National Forest27,643
West GirardShasta-Trinity National Forest37,516
References (14)
  1. Brown, Larry (U.S. Geological Survey, WRD). 2000. Review and annotation of fish watershed distribution maps. Review requested by Anthony E. Zammit, TNC. June 2000.
  2. Lee, D. S., C. R. Gilbert, C. H. Hocutt, R. E. Jenkins, D. E. McAllister, and J. R. Stauffer, Jr. 1980. Atlas of North American freshwater fishes. North Carolina State Museum of Natural History, Raleigh, North Carolina. i-x + 854 pp.
  3. Master, L. L. and A. L. Stock. 1998. Synoptic national assessment of comparative risks to biological diversity and landscape types: species distributions. Summary Report submitted to Environmental Protection Agency. The Nature Conservancy, Arlington, VA. 36 pp.
  4. Mayden, R. L., W. J. Rainboth, and D. G. Buth. 1991. Phylogenetic systematics of the cyprinid genera <i>Mylopharodon</i> and <i>Ptychocheilus</i>: comparative morphometry. Copeia 1991:819-834.
  5. Moyle, P. B. 1976a. Inland fishes of California. University of California Press, Berkeley, California. 405 pp.
  6. Moyle, P. B. 2002. Inland fishes of California. Revised and expanded. University of California Press, Berkeley. xv + 502 pp.
  7. Moyle, P. B., J. E. Williams, and E. D. Wikramanayake. 1989. Fish species of special concern of California. Final report submitted to California Dept. of Fish and Game, Inland Fisheries Division, Rancho Cordova. 222 pp.
  8. Nelson, J. S., E. J. Crossman, H. Espinosa-Perez, L. T. Findley, C. R. Gilbert, R. N. Lea, and J. D. Williams. 2004. Common and scientific names of fishes from the United States, Canada, and Mexico. American Fisheries Society, Special Publication 29, Bethesda, Maryland. 386 pp.
  9. Page, L. M., and B. M. Burr. 1991. A field guide to freshwater fishes: North America north of Mexico. Houghton Mifflin Company, Boston, Massachusetts. 432 pp.
  10. Page, L. M., and B. M. Burr. 2011. Peterson field guide to freshwater fishes of North America north of Mexico. Second edition. Houghton Mifflin Harcourt, Boston. xix + 663 pp.
  11. Page, L. M., H. Espinosa-Pérez, L. T. Findley, C. R. Gilbert, R. N. Lea, N. E. Mandrak, R. L. Mayden, and J. S. Nelson. 2013. Common and scientific names of fishes from the United States, Canada, and Mexico. Seventh edition. American Fisheries Society, Special Publication 34, Bethesda, Maryland.
  12. Page, L. M., K. E. Bemis, T. E. Dowling, H.S. Espinosa-Pérez, L.T. Findley, C. R. Gilbert, K. E. Hartel, R. N. Lea, N. E. Mandrak, M. A. Neigbors, J. J. Schmitter-Soto, and H. J. Walker, Jr. 2023. Common and scientific names of fishes from the United States, Canada, and Mexico. Eighth edition. American Fisheries Society (AFS), Special Publication 37, Bethesda, Maryland, 439 pp.
  13. Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea, and W.B. Scott. 1991. Common and scientific names of fishes from the United States and Canada. American Fisheries Society, Special Publication 20. 183 pp.
  14. State Natural Heritage Data Centers. 1996b. Aggregated element occurrence data from all U.S. state natural heritage programs, including the Tennessee Valley Authority, Navajo Nation and the District of Columbia: Export of freshwater fish and mussel records west of the Mississippi River in 1997. Science Division, The Nature Conservancy.