Dipodomys ingens

(Merriam, 1904)

Giant Kangaroo Rat

G1Critically Imperiled (G1G2) Found in 17 roadless areas NatureServe Explorer →
G1Critically ImperiledGlobal Rank
EndangeredIUCN
MediumThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.103814
Element CodeAMAFD03080
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNEndangered
Endemicendemic to a single state or province
KingdomAnimalia
PhylumCraniata
ClassMammalia
OrderRodentia
FamilyHeteromyidae
GenusDipodomys
Concept Reference
Wilson, D. E., and D. M. Reeder (editors). 1993. Mammal species of the world: a taxonomic and geographic reference. Second edition. Smithsonian Institution Press, Washington, DC. xviii + 1206 pp. Available online at: http://www.nmnh.si.edu/msw/.
Conservation Status
Review Date2014-09-23
Change Date2014-09-23
Edition Date2011-04-06
Edition AuthorsWilliams, D. F., and G. Hammerson
Threat ImpactMedium
Range Extent5000-20,000 square km (about 2000-8000 square miles)
Number of Occurrences6 - 20
Rank Reasons
Confined to southern California; most populations occupy relatively small areas of suitable habitat; habitat still being lost to various kinds of development; recent protection on the Carrizo Plain has alleviated threats somewhat; lack of appropriate habitat management on conservation lands may be resulting in habitat degradation in some areas.
Range Extent Comments
The range is confined to a narrow strip along the southwestern border of the San Joaquin Valley and a few nearby valleys to the west, including the Carrizo and Elkhorn plains and upper Cuyama Valley, with scattered colonies in the Ciervo, Kettleman, Panoche, and Turney Hills, and the Panoche Valley (Grinnell 1922, Hall 1981, Williams and Kilburn 1991, Williams et al. 1993). Historical range extended from Merced County south to the base of the Tehachapi Mountains in Kern County, and west to eastern San Luis Obispo County and extreme northern Santa Barbara County (Williams et al. 1993). Elevational range extends to about 868 meters (400-2,850 feet).

The population is currently fragmented into six major geographic units: 1) the Panoche region in western Fresno and eastern San Benito counties; 2) Kettleman Hills in Kings County; 3) San Juan Creek Valley in San Luis Obispo County; 4) western Kern County in the area of the Lokern, Elk Hills, and other uplands around McKittrick, Taft, and Maricopa; 5) Carrizo Plain in eastern San Luis Obispo County; and 6) Cuyama Valley in Santa Barbara and San Luis Obispo counties (see USFWS 1998). These major units are fragmented into more than 100 smaller populations, many of which are isolated by several miles of barriers such as steep terrain with plant communities unsuitable as habitat, or agricultural, industrial, or urban land without habitat for this species (USFWS 1998).
Occurrences Comments
The extant distribution includes more than 100 more or less distinct populations, but most of these represent fragments of a formerly more continuous distribution. There are six major geographic units, each of which could be regarded as a single occurrence or subpopulation.
Threat Impact Comments
The decline is largely a result of conversion of habitat to agricultural uses, combined with additional loss of habitat to industrial uses (e.g., petroleum exploration and extraction) and urbanization, and population reductions and extirpations from rodenticide use (aimed at California ground squirrel and historically at kangaroo rats) (Williams 1992, USFWS 1998). Possibly excessive livestock grazing may have contributed to the decline in some areas (livestock may crush near-surface burrows and compete for food with kangaroo rats) (see USFWS 1998). Between 1972 and 1980 most of the remaining habitat was converted from native vegetation to cultivated agricultural crops due in part to an abundance of irrigation water supplied by recently completed water delivery systems.

More recently, conversion of habitat to agricultural uses has slowed substantially, but urban and industrial developments, petroleum and mineral exploration and extraction, new energy and water conveyance facilities, and construction of communication and transportation infrastructures continue to destroy habitat and increase the threats to the species by reducing and further fragmenting populations (USFWS 1998). Habitat degradation due to lack of appropriate habitat management on conservation lands, especially lack of grazing or fire to control density of vegetation (including shrubs) may be a threat (Williams and Germano 1993).
Ecology & Habitat

Description

This is a very large, brownish kangaroo rat with a light brown tip on the tail; tail usually is less than 130 percent of the length of the head and body; hind foot has five toes; total length 311-348 mm (Jameson and Peeters 2004).

Diagnostic Characteristics

Other kangaroo rats in California have only four toes on the hind foot or usually have a head and body length of less than 130 mm (Jameson and Peeters 2004).

Habitat

Habitat consists of gently sloping and level piedmont plains and (formerly) areas supporting saltbush and perennial grasses; habitat now is dominated by introduced annuals, with many shrubs in some areas. The species occupies areas of sparse vegetative cover and well-drained soils and slope generally less than 9% (Williams and Kilburn 1991) (sometimes up to 22%; USFWS 1998), often in areas that are heavily grazed by cattle and sheep (Williams and Kilburn 1991). This kangaroo rat prefers semi-arid slopes at the head of draws in barren shrubless areas, with loose, easily diggable, sandy loam soils. When inactive, it occupies underground burrows. It is absent from areas continuously in dry-land cultivation and from irrigated fields but may recolonize fallow dry-land grain fields if there are colonies on uncultivated land nearby (Williams and Kilburn 1991). Habitats listed in order of decreasing favorability: 1) annual grassland association in areas with less than 5-6 inches annual rain, and level to gently sloping ground, 2) alkali desert scrub association in areas with less than 5-6 inches annual rain, sandy loam soils, and level to gently sloping ground, 3) friable soils of sand, loam, clay loam or gravelly in areas with the above charcateristics, and 4) slopes of 10-15 degrees with the above characteristics and located near colonies in more favorable habitats (D. Williams, pers. comm.). Young are born in burrows.

Ecology

Population density is 5-50 per ha (18-69/ha in the larger and denser colonies, Williams and Kilburn 1991). Home range is about 60-350 sq m; strict intrasexual avoidance indicated (basically solitary and territorial). Extant population sizes are small; from fewer than 10 to several hundred individuals (Braun 1985).

In spring, areas around occupied burrows have a more lush growth of herbaceous vegetation than do areas between burrow systems; this growth is eventually removed by grazing by livestock and/or kangaroo rats (Williams and Kilburn 1991).

Digging and feeding activity promote the establishment of exotic weeds, which in turn are a favored food source (Schiffman 1994).

Reproduction

Limited data indicate that the reproductive season may extend from January through May (Williams and Kilburn 1991) (February to June or perhaps later according to Biosystems Analysis 1989). Gestation lasts about 1 month. Litter size is 3-6 (Biosystems Analysis 1989); modal litter size probably is 4 (Williams and Kilburn 1991). Young are weaned at 4 weeks (Matthews and Moseley 1990). Flexible mating system (Hekkala, no date).
Terrestrial Habitats
Grassland/herbaceous
Other Nations (1)
United StatesN1
ProvinceRankNative
CaliforniaS2Yes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentLarge - smallExtreme or 71-100% pop. declineHigh (continuing)
1.1 - Housing & urban areasLarge - smallExtreme or 71-100% pop. declineHigh (continuing)
1.2 - Commercial & industrial areasLarge - smallExtreme or 71-100% pop. declineHigh (continuing)
2 - Agriculture & aquacultureRestricted - smallExtreme or 71-100% pop. declineHigh (continuing)
2.1 - Annual & perennial non-timber cropsRestricted - smallExtreme or 71-100% pop. declineHigh (continuing)
2.3 - Livestock farming & ranchingLarge - smallSerious - slightHigh (continuing)
3 - Energy production & miningLarge - smallExtreme - moderateHigh (continuing)
3.1 - Oil & gas drillingLarge - smallExtreme - moderateHigh (continuing)
4 - Transportation & service corridorsRestricted - smallExtreme - moderateHigh (continuing)
4.1 - Roads & railroadsRestricted - smallExtreme - moderateHigh (continuing)
4.2 - Utility & service linesRestricted - smallExtreme - moderateHigh (continuing)
7 - Natural system modificationsRestricted - smallExtreme - moderateHigh (continuing)
7.1 - Fire & fire suppressionRestricted - smallExtreme - moderateHigh (continuing)
8 - Invasive & other problematic species, genes & diseasesUnknownModerate - slightHigh (continuing)
8.1 - Invasive non-native/alien species/diseasesUnknownModerate - slightHigh (continuing)

Roadless Areas (17)
California (17)
AreaForestAcres
AntimonyLos Padres National Forest40,911
Big RocksLos Padres National Forest11,866
Black MountainLos Padres National Forest16,818
CuyamaLos Padres National Forest19,631
Fox MountainLos Padres National Forest52,072
Fox MountainLos Padres National Forest52,072
Garcia MountainLos Padres National Forest7,850
La BreaLos Padres National Forest14,031
La PanzaLos Padres National Forest4,954
Los Machos HillsLos Padres National Forest11,112
Lpoor CanyonLos Padres National Forest13,762
Lpoor CanyonLos Padres National Forest13,762
Machesna MountainLos Padres National Forest12,271
QuatalLos Padres National Forest7,253
Sawmill - BadlandsLos Padres National Forest51,362
Sespe - FrazierLos Padres National Forest106,910
Stanley MountainLos Padres National Forest14,674
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