Lithobates heckscheri

(Wright, 1924)

River Frog

G5Secure Found in 2 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Least concernIUCN
Identity
Unique IDELEMENT_GLOBAL.2.103815
Element CodeAAABH01120
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicendemic to a single nation
KingdomAnimalia
PhylumCraniata
ClassAmphibia
OrderAnura
FamilyRanidae
GenusLithobates
Synonyms
Aquarana heckscheri(Wright, 1924)Rana heckscheriWright, 1924
Other Common Names
river frog (EN)
Concept Reference
Frost, D. R. 1985. Amphibian species of the world. A taxonomic and geographical reference. Allen Press, Inc., and The Association of Systematics Collections, Lawrence, Kansas. v + 732 pp.
Taxonomic Comments
The generic taxonomy of American ranids is controversial, with three nomenclatural arrangements being consistent with current understandings of phylogeny: The single-genus arrangement of Yuan et al. (2016) which placed all Eurasian Rana and Pseudorana as well as all American ranids into Rana; the three-genus model of Che et al. (2007), largely in agreement with the earlier arrangement by Frost et al. (2006), which recognizes Pseudorana in Asia, Rana in Eurasia and western North America, and Lithobates in the Americas; and a seven-genus model of Dubois et al. (2021) which recognizes Pseudorana, Rana, and Liuhurana in Eurasia and Amerana (the Pacific Coast ranids of North America), Aquarana (for the bullfrogs and allies), Boreorana (a monotypic genus for Wood Frog, L. sylvaticus), and Lithobates (for the leopard frogs and allies). Here we follow Nicholson (2025) who defer from recognizing Amerana, Aquarana, or Boreorana at this time pending achievement of more phylogenetic stability, especially with respect to the position of the taxon sylvaticus which inconsistently is recovered as sister to the proposed Aquarana or as sister to Lithobates.
Conservation Status
Review Date2002-04-12
Change Date2001-11-21
Edition Date2002-04-12
Edition AuthorsHammerson, G.
Range Extent20,000-2,500,000 square km (about 8000-1,000,000 square miles)
Range Extent Comments
Coastal Plain from southern North Carolina (at least formerly) to northern Florida, west through extreme southern Alabama to southeastern Mississippi (Conant and Collins 1991). Most of the range is in northern Florida, southern Georgia, and southern South Carolina.
Ecology & Habitat

Habitat

Swamps along streams and edges of shallow ponds and impoundments (including beaver ponds). Eggs and larvae develop in permanent swamp waters.

Reproduction

Lays clutch of probably several thousand eggs April to August. Aquatic larval stage lasts 1-2 years.
Palustrine Habitats
FORESTED WETLAND
Other Nations (1)
United StatesN5
ProvinceRankNative
MississippiS1Yes
GeorgiaS5Yes
FloridaS3Yes
South CarolinaS4Yes
North CarolinaSXYes
AlabamaS1Yes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentHigh (continuing)
1.1 - Housing & urban areasHigh (continuing)
1.2 - Commercial & industrial areasHigh (continuing)
1.3 - Tourism & recreation areasHigh (continuing)
4 - Transportation & service corridorsHigh (continuing)
4.1 - Roads & railroadsHigh (continuing)

Roadless Areas (2)
Florida (2)
AreaForestAcres
Gum BayApalachicola National Forest11,645
SavannahApalachicola National Forest1,927
References (18)
  1. Bartlett, R. D., and P. P. Bartlett. 1999b. A field guide to Florida reptiles and amphibians. Gulf Publishing Company, Houston, Texas. xvi + 278 pp.
  2. Behler, J. L., and F. W. King. 1979. The Audubon Society field guide to North American reptiles and amphibians. Alfred A. Knopf, New York. 719 pp.
  3. Blackburn, L., P. Nanjappa, and M. J. Lannoo. 2001. An Atlas of the Distribution of U.S. Amphibians. Copyright, Ball State University, Muncie, Indiana, USA.
  4. Che, J., J. Pang, E. M. Zhao, M. Matsui, and Y. P. Zhang. 2007. Phylogenetic relationships of the Chinese brown frogs (genus <i>Rana</i>) inferred from partial mitochondrial 12S and 16S rRNA gene sequences. Zoological Science 24(1):71-80.
  5. Conant, R. 1975. A Field Guide to Reptiles and Amphibians of Eastern and Central North America. Second Edition. Houghton Mifflin Company, Boston, Massachusetts. xvii + 429 pp.
  6. Conant, R. and J. T. Collins. 1991. A field guide to reptiles and amphibians: eastern and central North America. Third edition. Houghton Mifflin Co., Boston, Massachusetts. 450 pp.
  7. Crother, B. I. (editor). 2017. Scientific and standard English names of amphibians and reptiles of North America north of Mexico, with comments regarding confidence in our understanding. 8th edition. SSAR Herpetological Circular 43:1-104. [Updates in SSAR North American Species Names Database at: https://ssarherps.org/cndb]
  8. Dubois, A., A. Ohler, and R. A. Pyron. 2021. New concepts and methods for phylogenetic taxonomy and nomenclature in zoology, exemplified by a new ranked cladonomy of recent amphibians (Lissamphibia). Megataxa 5:1-738.
  9. Frost, D. R. 1985. Amphibian species of the world. A taxonomic and geographical reference. Allen Press, Inc., and The Association of Systematics Collections, Lawrence, Kansas. v + 732 pp.
  10. Frost, D. R. 2010. Amphibian Species of the World: an Online Reference. Version 5.4 (8 April 2010). Electronic Database accessible at http://research.amnh.org/herpetology/amphibia/index.php. American Museum of Natural History, New York, USA.
  11. Frost, D.R. 2020. Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History, New York, USA. Online: http://research.amnh.org/herpetology/amphibia/index.html
  12. Frost, D. R. 2026. Amphibian Species of the World: an Online Reference. Version 6.2. Electronic Database accessible at https://amphibiansoftheworld.amnh.org/index.php. American Museum of Natural History, New York, USA. doi.org/10.5531/db.vz.0001
  13. Frost, D.R., T. Grant, J. Faivovich, R. Bain, A. Haas, C.F.B. Haddad, R.O. de Sa´, S.C. Donnellan, C.J. Raxworthy, M. Wilkinson, A. Channing, J.A. Campbell, B.L. Blotto, P. Moler, R.C. Drewes, R.A. Nussbaum, J.D. Lynch, D. Green, and W.C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1-370.
  14. Martof, B. S., W. M. Palmer, J. R. Bailey, and J. R. Harrison, III. 1980. Amphibians and reptiles of the Carolinas and Virginia. University of North Carolina Press, Chapel Hill, North Carolina. 264 pp.
  15. Mount, R. H. 1975. The reptiles and amphibians of Alabama. Auburn University Agricultural Experiment Station, Auburn, Alabama. vii + 347 pp.
  16. Nicholson, K. E. (ed.). 2025. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Ninth Edition. Society for the Study of Amphibians and Reptiles. 87 pp. Online database available at: https://cnah.org/SSARnames.aspx
  17. Sanders, A.E. 1984. <i>Rana heckscheri</i>. Catalogue of American Amphibians and Reptiles. 348:1-2.
  18. Yuan, Z.-Y, W.-W. Zhou, X. Chen, N.A. Poyarkov Jr, H.-M. Chen, N.-H. Jang-Liaw, W.-H. Chou, N.J. Matzke, K. Iizuka, M.-S. Min, S.L. Kuzmin, Y.-P. Zhang, D.C. Cannatella, D.M. Hillis, and J. Chesp. 2016. Spatiotemporal diversification of the true frogs (genus <i>Rana</i>): a historical framework for a widely studied group of model organisms. Systematic biology 65(5):824-842.