Occurrences CommentsIt is in every drainage (extirpated from Minnesota River) in most Minnesota Cos. (Sietman, 2003); incl. Red, Lake Superior, Lake of the Woods (Graf, 1997; Cvancara, 1970). In Illinois, it is in small/medium streams throughout (Cummings and Mayer, 1997; Schanzle and Cummings, 1991; Schanzle et al. 2004; Tiemann et al., 2005). Indiana: Blue (Sietman et al., 1995), Tippecanoe (Cummings and Berlocher, 1990), E Fork White (Harmon, 1992), Muscatatuck (Harmon, 1989), St. Joseph, Maumee and Eel (Pryor, 2005). In Ohio, it is throughout (Watters, 1992; 1995; Lyons et al., 2007; Grabarciewicz, 2008; Hoggarth et al., 2007; Watters et al., 2009). In West Virginia, it is in the Upper Ohio/Kanawha (Zeto et al., 1987) and Mud Rivers (Guyandotte drainage) (Schmidt and Zeto, 1986). In South Dakota it is in the Minnesota, upper/mid Big Sioux, James, Vermillion Rivers, and Lake Kampeska (Skadsen and Perkins, 2000; Backlund, 2000; Perkins and Backlund, 2003). In Montana, it is in most tribs. of the Missouri, Marais, Milk, Yellowstone, Little Missouri Rivers; less common in Tongue and Bighorn Rivers (Yellowstone basin), Battle and Beaver Creeks (Milk basin) (Gangloff and Gustafson, 2000; Stagliano, 2010). It is in Arkansas in the Poteau (Vaughn and Spooner, 2004), Cache (Christian et al., 2005), and mouth of White (Gordon, 1982). It is widespread and abundant in Wisconsin (Mathiak, 1979). In Mississippi, it is in the Mississippi River S, Big Black, and Yazoo drainages (Jones et al., 2005). It is rare in Louisiana, from Bayou Bartholomew, Tensas River (questionably) and nearby in W Mississippi (Vidrine, 1993). In Tennessee, it is only in Reelfoot Lake (nearly extirpated) and 1 Wolf trib. (Kesler and Manning, 1996) of the Mississippi (Parmalee and Bogan, 1998). It is statewide in Kentucky (Cicerello and Schuster, 2003); rarely in the Middle Green (Cochran and Layzer, 1993; Gordon, 1991), also S Fork Kentucky (Evans, 2008). It is widespread in Oklahoma in the Kiamichi, Blue, Clear Boggy and Little (Vaughn and Taylor, 1999; Vaughn, 2000) and Bird, Fourteenmile, Pryor and Big Cabin Creeks; Flint Creek; Haw Creek (LeFlore Co.); Glover Rivers and Pennington and Gates Creeks, Spring (Branson, 1966); Neosho, Mountain Fork (Spooner and Vaughn, 2007) and Blue Rivers and Honey and Sparrow Creek (Murray Co.); Poteau River; and Bird, Hominy and Salt Creeks (Branson, 1984). Kansas: Neosho, Marais des Cygnes, and Spring (Branson, 1966) basins, and (declined in Verdigris basin) still upper Fall and Caney Rivers but extirpated from Wakarusa (Tiemann, 2006), Republican, Solomon, Saline, Smoky Hill, Ninnescah, Walnut, and Cottonwood Rivers (Couch, 1997). Recently only dead shells in the Marais des Cygnes, Elk, and Fall Rivers, Kansas (Combes and Edds, 2005). In the Little Blue basin it is in the Kansas and Nebraska portions (Hoke, 2004). In the Big Blue system (SE Nebraska, NE Kansas) it was common as weathered valves and alive in W Fork Big Blue and a hole in the Big Blue mainstem, Nebraska (no recruitment) (Hoke, 2005). It is also known from the Clinton River drainage in Michigan (Trdan and Hoeh, 1993) and upper peninsula (Goodrich and Van der Schalie, 1939). It occurs in the Kalamazoo River (Mulcrone and Mehlne, 2001) and Lakes Michigan, Huron, and St. Clair drainages (Badra and Goforth, 2003), Michigan (Strayer, 1980; Trdan and Hoeh, 1993). Beetle (1989) lists Wyoming occurrences in Converse, Fremont, Natrona, and Platte Cos.; while Cvancara (2005) lists the Bighorn (Big Horn, Freemont Cos.), Tongue (Sheridan Co.), North Platte drainages (Natrona, Converse, Platte Cos.). In Canada it is widespread and abundant in Manitoba (Assiniboine- Watson, 2000; Pip, 2006), Ontario (Metcalfe-Smith et al., 2003), Quebec, Saskatchewan, and less abundant, but widespread in Northwest Territories, and Alberta (most SE of the Cordillera); and undetermined in Nunavut (Metcalfe-Smith and Cudmore-Vokey, 2004); recently Petitot River in British Columbia (BC CDC, pers. comm., 2010).