Artemisiospiza belli

(Cassin, 1850)

Bell's Sparrow

G5Secure Found in 27 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Least concernIUCN
Identity
Unique IDELEMENT_GLOBAL.2.902234
Element CodeABPBX97020
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassAves
OrderPasseriformes
FamilyPasserellidae
GenusArtemisiospiza
Synonyms
Amphispiza belli(Cassin, 1850)
Other Common Names
Bell's sparrow (EN) Bruant de Bell (FR)
Concept Reference
American Ornithologists' Union (AOU). Chesser, T.R., R.C. Banks, F.K. Barker, C.Cicero, J.L. Dunn, A.W. Kratter, I.J. Lovette, P.C. Rasmussen, J.V. Remsen Jr., J.D.Rising, D.F. Stotz and K.Winker. 2013. Fifty-Fourth Supplement to the American Ornithologists' Union Check-list of North American Birds. Auk 130(3):558-571.
Taxonomic Comments
Artemisiospiza nevadensis was formerly considered conspecific with A. belli, but treated as a separate species on the basis of differences in mitochondrial DNA, morphology, and ecology, and limited gene flow at the contact zone in eastern California (Cicero and Johnson 2007, Cicero and Koo 2012) (AOU 2013).

Populations of A. b. canescens of the San Joaquin Valley and Mojave Desert differ in morphology and ecology from belli and may represent a distinct species. Analyses of mtDNA indicate that Mojave Desert populations of canescens are distinctive, whereas canescens from the San Joaquin Valley share haplotypes with coastal belli (Cicero and Koo 2012) (AOU 2013).

Formerly placed in the genus Amphispiza, but genetic data (Klicka and Spellman 2007, DaCosta et al. 2009) indicate that the new genera are not closely related (AOU 2012). This was formerly proposed in Klicka and Banks (2011).
Conservation Status
Review Date1999-11-20
Change Date1996-12-04
Edition Date1999-11-20
Edition AuthorsRevisions by M. KOENEN and D.W. MEHLMAN.
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences81 to >300
Rank Reasons
Large range in the western U.S. and Mexico; fairly common and stable in many areas, with local declines.
Range Extent Comments
BREEDING: central Washington, eastern Oregon, southern Idaho, southwestern Wyoming, and northwestern Colorado south to southern California, central Baja California, southern Nevada, southwestern Utah, northeastern Arizona, and northwestern New Mexico (AOU 1983, Martin and Carlson 1998). NON-BREEDING: central California, central Nevada, southwestern Utah, northern Arizona, and central New Mexico south to central Baja California, northwestern mainland of Mexico, and western Texas (AOU 1983, Martin and Carlson 1998). Sedentary subspecies BELLI: foothills of the West coast (northern California to northwestern Baja California) and the western slope of the central Sierra Nevada in California (Johnson and Marten 1992). Subspecies CANESCENS: breeds in the San Joaquin Valley and northern Mohave Desert in California and extreme western Nevada, winters in the southwestern U.S. (Johnson and Marten 1992). Subspecies NEVADENSIS: breeds from central interior Washington eastward to southwestern Wyoming and northwestern Colorado, south to east-central California, central Nevada, northeastern Arizona, and northwestern New Mexico; winters in the southwestern U.S. and northern Mexico (Johnson and Marten 1992).
Occurrences Comments
No exact figures available.
Threat Impact Comments
HABITAT LOSS, FRAGMENTATION: Sagebrush shrublands are vulnerable to a number of activities that reduce or fragment sagebrush habitat, including land conversion to tilled agriculture, urban and suburban development, and road and power-line rights of way. Range improvement programs remove sagebrush by burning, herbicide application, and mechanical treatment, replacing sagebrush with annual grassland to promote forage for livestock. West of the Rocky Mountains, in the Great Basin and interior Columbia Basin, there is concern that livestock grazing can damage soil, specifically the delicate cryptogamic layer, to the extent that vegetative succession is altered and recovery is hampered (Saab et al. 1995).

INVASIVE GRASSES: Cheatgrass readily invades disturbed sites, and has come to dominate the grass-forb community of more than half the sagebrush region in the West, replacing native bunchgrasses (Rich 1996). Crested wheatgrass and other non-native annuals have also fundamentally altered the grass-forb community in many areas of sagebrush shrub-steppe.

FIRE: Cheatgrass has altered the natural fire regime in the western range, increasing the frequency, intensity, and size of range fires. Fire kills sagebrush and where non-native grasses dominate, the landscape can be converted to annual grassland as the fire cycle escalates, removing habitat for sage sparrow (Paige and Ritter 1998).

BROOD PARASITISM: An occasional host for brown-headed cowbird (Molothrus ater), and may abandon nest (e.g., see Reynolds 1981). Prior to European-American settlement, was probably largely isolated from cowbird brood parasitism, but is now vulnerable where the presence of livestock, land conversion to agriculture, and fragmentation of shrublands creates a contact zone between the species (Rich 1978).

INTRODUCED ANIMALS: Pigs, goats, and other grazing animals introduced onto San Clemente Island, California, reduced habitat and local numbers of the subspecies declined (Everatt et al. 1994). Feral cats near human habitations may increase predation (Martin and Carlson 1998).

PREDATION: In Oregon, predation by Townsend ground squirrel (Spermophilus townsendi) affected reproductive success when squirrel densities were high; populations in southeastern Washington and northern Nevada incurred high rates of nest predation, probably mainly by gopher snakes (Pituophis melanoleucus) (Rotenberry and Wiens 1989). Loggerhead shrikes (Lanius ludovicianus) prey on both adults and altricial young in nest, and can significantly reduce nest production (Reynolds 1979). Other predators include common raven (Corvus corax) and merlin (Falco columbarius; Martin and Carlson 1998).
Ecology & Habitat

Description

Gray-brown head, white eye ring, white lore spot or eyebrow, and broad white whisker stripe above a dark whisker stripe; back is buffy brown with dusky streaks; two pale wing bars (not conspicuous); underparts are white, with a central dark spot and dusky streaks on the sides; juveniles are duller and more heavily streaked, and the lore spot sometimes is indistinct (NGS 1983). Subspecies BELLI of western California is smaller and darker than are subspecies CANESCENS and NEVADENSIS (Johnson and Marten 1992).

Diagnostic Characteristics

Differs from the five-striped sparrow (AIMOPHILA QUINQUESTRIATA) in lacking a gray breast and sides (adults) and by the presence of streaks in the juvenile plumage. Differs from the lark sparrow (CHONDESTES GRAMMACUS) by lacking the bold head pattern of rusty, black, and white; and by lacking white corners on the end of the tail. Lacks the gray and rufous head and unstreaked sides of the American tree sparrow (SPIZELLA ARBOREA).

Habitat

BREEDING: Found from sea level to 2000 meters (Rising 1996); strongly associated with sagebrush for breeding. Also found in salt-bush brushland, shadscale, antelope brush, rabbitbrush, black greasewood (Colorado), mesquite, and chaparral (California; AOU 1998; Green and Smith 1981; Martin and Carlson 1998; Paige and Ritter 1998; Reynolds 1981). Prefers semi-open habitats, shrubs 1-2 meters tall (Martin and Carlson 1998). Habitat structure (vertical structure, shrub density, and habitat patchiness) is important to habitat selection (Martin and Carlson 1998). Positively correlated with big sagebrush (Artemisia tridentata), shrub cover, bare ground, above-average shrub height, and horizontal patchiness; negatively correlated with grass cover (Rotenberry and Wiens 1980; Wiens and Rotenberry 1981; Larson and Bock 1984).

In the northern Great Basin, this species is associated with low and tall sagebrush/bunchgrass, juniper/sagebrush, mountain mahogany/shrub, and aspen/sagebrush/bunchgrass communities for breeding and foraging (Maser et al. 1984). In Idaho, it is found in sagebrush of 11 to 14 percent cover (Rich 1980). Martin and Carlson (1998) report preference for evenly spaced shrubs; other authors (Rotenberry and Wiens 1980; Peterson and Best 1985) report association where sagebrush is clumped or patchy.

In Nevada, sparrows are most abundant in sagebrush habitat, but they also breed in salt desert scrub more frequently than other sagebrush "obligate" birds. Greasewood may also be used as a breeding substrate with some frequency, although existing evidence is ambiguous (GBBO 2010). Some Sage Sparrows winter in southern Nevada, usually in sagebrush or Mojave scrub shrublands, but also in honey mesquite stands (GBBO 2010). In the Great Basin Bird Observatory study in Nevada, Sage Sparrows were strongly associated with greater shrub cover and the absence of trees; they were negatively associated with greater herbaceous cover, shrub height was not a strong predictor, and there was no relationship to proximity of water (GBBO 2010).

Subspecies belli: chaparral dominated by chamise and/or California sagebrush (Johnson and Marten 1992). Subspecies canescens: breeds in desert scrub where Atriplex is prevalent (Johnson and Marten 1992). Subspecies nevadensis: breeds in brushland dominated by big sagebrush or sagebrush-saltbush (Johnson and Marten 1992). Subspecies clementeae: nests in boxthorn shrubs interspersed by cactus (Willey 1997).

Nests on the ground or in a shrub, up to about one meter above ground (Terres 1980). In the Great Basin, usually nests in living sagebrush where cover is sparse but shrubs are clumped; avoids southwestern side of plant (Petersen and Best 1985). Placement may be related to density of vegetative cover over the nest, as will nest higher in a taller shrub (Rich 1980).

NON-BREEDING: In migration and winter also in arid plains with sparse bushes, grasslands and open situations with scattered brush, mesquite, and riparian scrub; preferring to feed near woody cover (Martin and Carlson 1998; Meents et al. 1982; Repasky and Schluter 1994). Flocks in Mojave Desert appear to follow water courses (Eichinger and Moriarty 1985). Wintering birds in honey mesquite of lower Colorado River select areas of higher inkweed (Suaeda torreyana) density (Meents et al. 1982). Some Sage Sparrows winter in southern Nevada, usually in sagebrush or Mojave scrub shrublands, but also in honey mesquite stands (GBBO 2010).

Ecology

Breeding territory size usually averages about 1.5-3 hectares (Wiens et al. 1985). Territory size can vary with plant community composition and structure, increasing with horizontal patchiness (see Wiens et al. 1985); also positively correlated with reproductive success (Peterson and Best 1987b). In Great Basin, density usually 50-200 per square kilometer (Rotenberry and Wiens 1989).

Reproduction

Clutch size usually is three to four, sometimes five. Incubation lasts about 13 days. Nestlings are altricial. Individual females produce one to three broods annually. Reproductive success is greater in wetter years (Rotenberry and Wiens 1991).
Terrestrial Habitats
Shrubland/chaparralDesert
Other Nations (1)
United StatesN5
ProvinceRankNative
CaliforniaSNRB,SNRNYes
NevadaS3Yes
ArizonaS3NYes
Roadless Areas (27)
California (27)
AreaForestAcres
AntimonyLos Padres National Forest40,911
Bear MountainLos Padres National Forest913
Black ButteLos Padres National Forest5,116
Cahuilla MountainSan Bernardino National Forest6,952
City CreekSan Bernardino National Forest9,997
Cucamonga BSan Bernardino National Forest11,933
CuyamaLos Padres National Forest19,631
Deer MountainMendocino National Forest11,716
Dry LakesLos Padres National Forest17,043
Eagle PeakCleveland National Forest6,481
Fox MountainLos Padres National Forest52,072
Granite PeakSan Bernardino National Forest450
GrindstoneMendocino National Forest26,031
Hixon FlatSan Bernardino National Forest8,095
Lpoor CanyonLos Padres National Forest13,762
MatilijaLos Padres National Forest5,218
No NameCleveland National Forest4,897
Pleasant ViewAngeles National Forest26,395
Rouse HillSan Bernardino National Forest13,745
San DimasAngeles National Forest7,160
San SevaineSan Bernardino National Forest6,866
Sawmill - BadlandsLos Padres National Forest51,362
ScodiesSequoia National Forest725
Sespe - FrazierAngeles National Forest4,254
South SierraInyo National Forest41,853
South SierraSequoia National Forest8,008
Wonoga Pk.Inyo National Forest11,272
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