Myotis evotis

(H. Allen, 1864)

Long-eared Myotis

G5Secure Found in 30 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Least concernIUCN
LowThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.798516
Element CodeAMACC01070
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassMammalia
OrderChiroptera
FamilyVespertilionidae
GenusMyotis
Other Common Names
Chauve-souris à longues oreilles (FR) Long-eared Bat (EN) long-eared bat (EN) Un Murciélago (ES)
Concept Reference
Wilson, D. E., and D. M. Reeder (editors). 2005. Mammal species of the world: a taxonomic and geographic reference. Third edition. The Johns Hopkins University Press, Baltimore. Two volumes. 2,142 pp. [As modified by ASM the Mammal Diversity Database (MDD) at https://www.mammaldiversity.org/index.html]
Taxonomic Comments
This taxonomic concept follows The American Society of Mammalogists (ASM) and Bats of the World (Simmon and Cirranello 2025) and retains M. keenii and M. evotis as distinct species. This status was re-affirmed by the Global Bat Taxonomy Working Group (Upham et al. 2022). There remains debate over the status of keenii (see Nagorsen and Brigham 1993, Van Zyll de Jong and Nagorsen 1994, Nagorsen 2002, Lung et al. 2013, Lausen et al. 2019, Morales et al. 2021, and Lausen et al. 2021). According to ASM, "The burden of proof is on overturning the species-level status of Myotis keenii, which has been recognized consistently by taxonomic authorities since 1979 and is thus the prevailing usage in need of refutation."

Population genetic analyses by Lausen et al. (2019) revealed four well-defined groups (species): M. septentrionalis, M. thysanodes, M. lucifugus, and a single group comprising M. keenii and M. evotis. They conclude that M. keenii does not qualify as a genetic or biological species and should be synonymized with M. evotis. Morales et al. (2021) re-examined the data of Lausen et al. (2019) and disputed this conclusion, however, the rebuttal by Lausen et al. (2021), confirms the data were accurate.

M. milleri in Baja California was treated as a distinct species in Wilson and Reeder (1993) but was included in M. evotis by Simmons (in Wilson and Reeder 2005).
Conservation Status
Rank Method Rank calculation - Biotics v2
Review Date2021-04-30
Change Date2015-03-31
Edition Date2021-04-30
Edition AuthorsHammerson, G. (2015); rev. S. Cannings (2021)
Threat ImpactLow
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences81 to >300
Rank Reasons
Widespread distribution in western North America; many occurrences in wide range of habitats; occurs regularly in low numbers across range; uses wide range of often plentiful roost sites; trend uncertain but probably relatively stable or slowly declining; no known major threats, but range, habitats, and threats during winter are poorly known.
Range Extent Comments
Range includes western North America, from central British Columbia (and probably southern Yukon), southwestern Northwest Territories, and southern Saskatchewan south along the Pacific Coast to Baja California, east through Montana and Idaho to the western Dakotas, and from Nevada, Utah, Wyoming and Colorado to New Mexico and Arizona (Manning and Jones 1989, Adams 2003, Lausen 2006, Reid 2006, Slough pers. comm.). Elevational range extends from near sea level along the Pacific Coast to around 2,900 meters in Utah and Wyoming (Manning and Jones 1989, Adams 2003).
Occurrences Comments
The number of distinct occurrences has not been determined using standardized/meaningful criteria, but the species is represented by a large number of collection/observation sites and locations (as defined by IUCN). As evidence of this, Verts and Carraway (1998) mapped more than 100 collection sites in Oregon, Hoffmeister (1986) mapped 52 localities in Arizona, and Nagorsen and Brigham (1993) mapped approximately 49 collection sites in British Columbia.
Threat Impact Comments
Overall, no major threats are known. Locally, these bats may be affected to some degree by a multitude of factors, including the closure of abandoned (unsurveyed) mines, recreational caving, some forest management practices, and activities (such as highway construction, water impoundments, blasting of cliffs for avalanche control) that impact cliff faces or rock outcrops (Bogan et al. 2005). Broadcast application of pesticides to combat insect pests potentially has a detrimental impact on bat food resources or on the bats themselves, but population impacts are uncertain. Toxic impoundments associated with resource extraction are a potential source of bat mortality or contamination (Buseck and Keinath 2004).

Forest management practices that result in reduced availability of roost sites in snags and trees with loose bark may have a negative effect on these bats. Roosts under exfoliating bark may be relatively short-lived resources (Vonhof and Barclay 1996, New Mexico Department of Game and Fish 1997). However, stumps (Vonhof and Barclay 1997, Waldien et al. 2000) and rock crevices (if available) (Snider et al. 2013) may provide numerous suitable alternate roosts.

A single Myotis evotis was found dead with white-nose syndrome in Washington State (Washington Department of Fish and WIldlife 2019). At present, this threat is considered Low, but could increase if the disease spreads more widely in the west.

Increased incidence of wildfires might negatively affect bat food resources or degrade local environmental conditions, but substantial populations of M. evotis can persist even in extensively burned areas where suitable rock-crevice roosts are available (Snider et al. 2013).

This species is subject to mortality from turbines at wind energy facilities. Arnett and Baerwald (2013) estimated that approximately 3,730-7,330 individuals were killed by turbines in the United States and Canada during the period 2000-2011. Given the ongoing increase in turbine installation, this mortality may increase significantly during the foreseeable future. Although the current impact of turbine-associated mortality on the overall M. evotis population is probably relatively small, the reproductive rate for this species is low, and the ability of regional populations to sustain current and future levels of impact is uncertain.

The impact of potential threats associated with pollution and climate change are unknown.
Ecology & Habitat

Habitat

These bats occupy a diverse array of habitats, including lowland, montane, and subalpine woodlands, forests, shrublands, and meadows, wooded stream courses, and areas over water bodies (Findley et al. 1975, Hoffmeister 1986, Manning and Jones 1989, Armstrong et al. 1994, Oliver 2000, Adams 2003). Within a particular region, the range of occupied habitats may be more restricted. Daytime roosts are in buildings, railroad trestles, snags and hollow trees, spaces behind loose bark of trees or stumps, mines, caves, rock crevices (including those on the ground), erosional cavities and channels in the ground, and similar sites (Barbour and Davis 1969; Vonhof and Barclay 1996, 1997; Oliver 2000; Waldien et al. 2000; Rancourt et al. 2005; Solick and Barclay 2006; Nixon et al. 2009; Snider et al. 2013). In southwestern Colorado, researchers found 33 roosts of reproductive females in rock crevices and 1 in a juniper snag; the bats often switched roosts (Snider et al. 2013). Frequent use of rock crevices also has been documented in Washington (Rancourt et al. 2005) and Alberta (Solick and Barclay 2006). Frequent roost switching (within small roosting home-ranges) occurred in a ground-roosting population in Alberta (Nixon et al. 2009). Small maternity colonies of 12-30 individuals have been found in buildings in British Columbia (Cowan and Guiguet 1965), and a group of adults and young were found in an uninhabited ranch house in Colorado (Barbour and Davis 1969). In northeastern Washington, roosts of reproductive females were in crevices in small basalt rock formations (Rancourt et al. 2005). See Vonhof and Barclay (1996, 1997) for information on characteristics of roosts in trees and stumps in British Columbia. Hibernation sites are poorly known.

Ecology

Nonreproductive females and males generally roost singly or in small groups (Rancourt et al. 2005, Solick and Barclay 2006).

Reproduction

Births have been recorded in June-July. Litter size is 1.Young and lactating females were recorded in late July in New Mexico. Female and newborn young were recorded in late June in California. Male young-of-year about adult size were observed in early August in South Dakota. Maximum longevity is at least 22 years (Tuttle and Stevenson 1982). Reproductive females form small maternity colonies (generally fewer than 30 adults). In southwestern Colorado, reproductive females roosted alone or in small groups of 2-3 individuals (Snider et al. 2013).
Terrestrial Habitats
Forest/WoodlandForest - HardwoodForest - ConiferForest - MixedWoodland - HardwoodWoodland - ConiferWoodland - MixedShrubland/chaparralGrassland/herbaceousCliffUrban/edificarian
Palustrine Habitats
Riparian
Other Nations (2)
United StatesN4
ProvinceRankNative
ArizonaS3Yes
ColoradoS4Yes
South DakotaS1Yes
Navajo NationS3Yes
WyomingS4BYes
NevadaS3Yes
UtahS3Yes
MontanaS3Yes
IdahoS3Yes
OregonS4Yes
North DakotaSUYes
CaliforniaS3Yes
WashingtonS4Yes
CanadaNNR
ProvinceRankNative
Northwest TerritoriesSUYes
British ColumbiaSNRYes
AlbertaS3Yes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentNegligible (<1%)High (continuing)
2 - Agriculture & aquacultureLarge - smallNegligible or <1% pop. declineHigh (continuing)
2.3 - Livestock farming & ranchingHigh (continuing)
3 - Energy production & miningRestricted - smallModerate - slightHigh (continuing)
3.2 - Mining & quarryingNegligible (<1%)Extreme or 71-100% pop. declineHigh (continuing)
3.3 - Renewable energyRestricted - smallModerate - slightHigh (continuing)
4 - Transportation & service corridorsNegligible (<1%)Negligible or <1% pop. declineHigh (continuing)
5 - Biological resource useSmall (1-10%)Moderate - slightHigh (continuing)
5.3 - Logging & wood harvestingHigh (continuing)
6 - Human intrusions & disturbanceSmall (1-10%)Moderate - slightHigh (continuing)
6.1 - Recreational activitiesSmall (1-10%)Moderate - slightHigh (continuing)
7 - Natural system modificationsNegligible (<1%)Serious - slightHigh (continuing)
7.1 - Fire & fire suppressionHigh (continuing)
7.2 - Dams & water management/useModerate (short-term)
8 - Invasive & other problematic species, genes & diseasesSmall (1-10%)Extreme or 71-100% pop. declineHigh (continuing)
9 - PollutionUnknownUnknownHigh (continuing)
10 - Geological eventsNegligible (<1%)
11 - Climate change & severe weatherPervasive - largeUnknownHigh (continuing)
11.1 - Habitat shifting & alterationHigh (continuing)
11.2 - DroughtsHigh (continuing)

Roadless Areas (30)
California (16)
AreaForestAcres
AgnewSequoia National Forest9,561
Barker ValleyCleveland National Forest11,940
Bucks LakePlumas National Forest680
Chips CreekLassen National Forest29,089
Chips CreekPlumas National Forest12,940
Dinkey LakesSierra National Forest34,171
Duncan CanyonTahoe National Forest8,621
Hoover - NorthHumboldt-Toiyabe National Forest1,574
HortonInyo National Forest5,717
Iceberg - Mill CreekHumboldt-Toiyabe National Forest26,988
Kings RiverSierra National Forest52,999
Mill PeakSan Bernardino National Forest7,884
MonarchSierra National Forest697
MosesSequoia National Forest22,077
Mt. Shasta BShasta-Trinity National Forest2,809
North Fork American RiverTahoe National Forest38,495
Idaho (1)
AreaForestAcres
SeceshPayette National Forest248,088
Montana (3)
AreaForestAcres
QuiggLolo National Forest67,267
Silver KingLolo National Forest12,816
Silver KingBeaverhead-Deerlodge National Forest64,289
Nevada (2)
AreaForestAcres
Alta T - NorthHumboldt-Toiyabe National Forest9,325
Table Mtn. - EastHumboldt-Toiyabe National Forest87,789
New Mexico (1)
AreaForestAcres
Frisco BoxGila National Forest38,979
Oregon (2)
AreaForestAcres
Drift CreekSiuslaw National Forest6,333
LakeMt. Hood National Forest1,327
Utah (2)
AreaForestAcres
Clear CreekSawtooth National Forest7,188
Mt. AireWasatch-Cache National Forest9,681
Wyoming (3)
AreaForestAcres
Lake Alice - Commissary RidgeBridger-Teton National Forest166,707
Pacific Creek - Blackrock CreekBridger-Teton National Forest24,658
Spread Creek - Gros Ventre RiverBridger-Teton National Forest166,097
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