Myotis occultus

Hollister, 1909

Arizona Myotis

G4Apparently Secure (G4G5) Found in 1 roadless area NatureServe Explorer →
G4Apparently SecureGlobal Rank
Least concernIUCN
Medium - lowThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.638468
Element CodeAMACC01160
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicendemic to a single nation
KingdomAnimalia
PhylumCraniata
ClassMammalia
OrderChiroptera
FamilyVespertilionidae
GenusMyotis
Synonyms
Myotis lucifugus occultusHollister, 1909
Other Common Names
Arizona myotis (EN) Un Murciélago (ES)
Concept Reference
Piaggio, A. J., E. W. Valdez, M. A. Bogan, and G. S. Spicer. 2002. Systematics of Myotis occultus (Chiroptera: Vespertilionidae) inferred from sequences of two mitochondrial genes. Journal of Mammalogy 83:386-395.
Taxonomic Comments
The bats in the Myotis lucifugus-M. occultus-M. fortidens complex have long been of debatable taxonomic status (Findley and Jones 1967). Myotis occultus was formerly included in M. lucifugus (Jones et al. 1992; Koopman, in Wilson and Reeder 1993), but now regarded as a distinct species. Allozyme data suggested that the two are conspecific (Valdez et al. 1999), but mitochondrial DNA and morphological evidence suggest that M. occultus is a specifically distinct, monophyletic lineage (Piaggio et al. 2002).
Conservation Status
Rank MethodLegacy Rank calculation - Excel v3.1x
Review Date2015-04-03
Change Date2015-04-03
Edition Date2015-04-03
Edition AuthorsHammerson, G.
Threat ImpactMedium - low
Range Extent200,000-2,500,000 square km (about 80,000-1,000,000 square miles)
Rank Reasons
Moderately large range in the American Southwest and northern and central Mexico; many roost sites in snags, also uses buildings and bridges; presumed large population size; vulnerable to various forms of habitat loss and degradation, especially those that affect roosts; better information is needed on current abundance, trend, and threats; some cave-hibernating popuations may be vulnerable to declines from rapidly spreading white-nose syndrome.
Range Extent Comments
The range encompasses northern and central Mexico, southeastern California, Arizona (Hoffmeister 1986, Arizona Game and Fish Department 2003), New Mexico (except perhaps extreme northeast; Frey 2004), southern Colorado, and perhaps southern Utah and western Texas (Piaggio et al. 2002; Valdez, pers. comm.). In Arizona, this bat is distributed predominantly in the highlands and upper stream reaches (Piaggio et al. 2002). In New Mexico, it is known from low-elevation riparian areas in the Rio Grande Valley and montane highlands. In California, it occurs only along the Colorado River lowlands and in the adjacent desert mountain ranges. The range meets that of M. lucifugus in central Colorado; that species occupies the northern third of the state (Valdez, pers. comm.). The only record from Texas is an unusual, perhaps intermediate specimen collected in 1893 near Fort Hancock in Hudspeth County; it is doubtful that a resident population exists in Texas (Piaggio et al. 2002, Schmidly 2004, Ammerman et al. 2012). Range limits in Mexico are uncertain; the species has been recorded from several localities in Chihuahua and a disjunct site near Texcoco in the Distrito Federal (Piaggio et al. 2002; Valdez, pers. comm.). In Arizona, this bat is most common at elevations of 1,830-2,806 meters, but it also occurs much lower along rivers in desert regions (Arizona Game and Fish Department 2003).
Occurrences Comments
The number of distinct occurrences has not been determined using standardized criteria, but the species is represented by a fairly large number of collection/observation sites and locations (as defined by IUCN).
Threat Impact Comments
Without citing specific examples or sources, Schmidly (2004) and Ammerman et al. (2012) reported that this species has declined sharply in much of its historical range in California as a result of pesticide use, control measures in nursery colonies, and disturbance at hibernation sites. However, the species historically occurred in California only along the lower Colorado River, and the species recently has been found there in riparian restoration sites (Calvert and Neiswenter 2012).

Cave/mine-roosting populations are vulnerable to disturbance (e.g., human intrusions) and to roosting habitat loss/degradation from mine closures or renewed mining.

Tree-roosting populations (e.g., Rabe et al. 1998) are vulnerable to factors that reduce the availability of roost sites (e.g., forest management practices that reduce the availability of snags, illegal firewood harvest, increased incidence of forest fires).

This species often is associated with large bodies of water that are unlikely to be greatly affected by water management.

Livestock grazing may potentially degrade bat foraging habitat by reducing prey availability, but the actual impact of grazing on populations of this species is uncertain.

Currently, this species is not known to incur significant mortality from turbines at wind energy facilities (Arnett and Baerwald 2013).

As of early 2015, this species was not known to be affected by white-nose syndrome, However, given its close (possibly conspecific) relationship with M. lucifugus (millions of which have been killed by WNS), certain cave/mine-hibernating populations of occultus may be regarded as potentially vulnerable to this rapidly spreading disease.
Ecology & Habitat

Description

Medium sized Myotis (total length = 80.0-97.0 mm [3.2-3.88 inches] and forearm length = 36.0-41.0 mm [1.44-1.64 inches]) with sleek glossy fur. Small ears (11.0-16.0 mm [0.44-0.64 inches]) and large feet (8.0-11.0 mm [0.32-0.44 inches]) are characteristic. Long hairs occur on the toes and extend beyond the tips of the claws. Color often bright, generally tawny, ochraceous, pale tan, or reddish-brown to dark brown. [From Arizona Game and Fish Department Heritage Data Management System]

Diagnostic Characteristics

This is the only long-footed (i.e. hind foot length >8.0 mm [0.32 inches]) Myotis in Arizona with a gradually sloping forehead and the only Myotis in Arizona with only 1 small upper premolar behind the canine. In the rare individual with 2, it is on 1 side only or 1 is crowded out of alignment. [From Arizona Game and Fish Department Heritage Data Management System]

The genus Myotis is distinguished from other bat species in Arizona by lack of a nose-leaf, or enlarged facial glands, or a tail extending beyond the tail membrane, or fur on the tail membrane. Myotis are initially identified by their uniform shades of brown and by their straight and relatively narrow tragus with a pointed tip. When compared to other Myotis, the lack of a keeled calcar distinguishes M. occultus from M. californicus, M. ciliolabrum, and M. volans. Shorter ears (11-16 mm) distinguish M. occultus from M. evotis (20-24 mm), M. auriculus (19-21 mm) and usually from M. thysanodes (12-19 mm). [From Arizona Game and Fish Department Heritage Data Management System]

Myotis occultus may also be distinguished from M. thysanodes by the lack of a macroscopic fringe of hairs on trailing edge of the tail membrane in M. occultus. Lack of bare spot between scapulae and lack of grayish back distinguish M. occultus from M. velifer. Darker ears and longer forearm (36.0-41.0 mm [1.44-1.64 inches]), and a glossier coat distinguish M. occultus from M. yumanensis which usually has light-colored ears, a shorter forearm (31.0-36.0 mm [1.24-1.44 inches]), and a dull coat. [From Arizona Game and Fish Department Heritage Data Management System]

Characters that are potentially useful for differentiating M. occultus and M. lucifugus carissima include the number of upper premolars (complete set in lucifugus), length of maxillary toothrow, presence or absence of sagittal crest (present in occultus, not in lucifugus), and pelage coloration (sometimes reddish in occultus, glossy brown in lucifugus) (Findley and Jones 1967), and rostrum length and slope of forehead (Hoffmeister 1986). However, morphologically intermediate specimens have been found, and at least some of these group genetically with M. occultus (Piaggio et al. 2002).

Habitat

Habitats in Arizona include ponderosa pine and oak-pine woodland near water; this bat is also known from wooded riparian areas in desert areas (Hoffmeister 1986, Morrell et al. 1999, Calvert and Neiswenter 2012). In New Mexico, the species is usually associated with large bodies of water without respect to associated vegetation type (Findley et al. 1975); it is known from low elevation riparian areas in the Rio Grande Valley and montane highlands (Piaggio et al. 2002).

Maternity colonies have been found in buildings (e.g., the attics of abandoned houses; Hoffmeister 1986; Chung-MacCoubrey 1999), in crevices of highway bridges (Stager 1943, Arizona Game and Fish Department 2003, Geluso and Mink 2009), and in snags (spaces under exfoliating ponderosa pine bark) (Rabe et al. 1998). In the valley of the Rio Grande in New Mexico, roosts were found in crevices among timbers of 9 bridges, 2 of which served as maternity roosts (Geluso and Mink 2009).

Winter habitat is poorly known. A few individuals have been found hibernating in mines in California and Sonora (Howell 1989). Hibernation also may occur in tree-hollow roosts in areas where the species occurs in summer (Ammerman et al. 2012).

Reproduction

In Arizona, births occur in May-June (Hoffmeister 1986, Calvert and Neiswenter 2012).
Terrestrial Habitats
Forest/WoodlandWoodland - HardwoodWoodland - ConiferWoodland - MixedDesertUrban/edificarian
Palustrine Habitats
RiparianAerial
Other Nations (1)
United StatesN4
ProvinceRankNative
New MexicoS4Yes
ColoradoSNRYes
CaliforniaS1Yes
ArizonaS3Yes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentNegligible (<1%)High (continuing)
2 - Agriculture & aquacultureLarge (31-70%)UnknownHigh (continuing)
2.1 - Annual & perennial non-timber cropsHigh (continuing)
2.3 - Livestock farming & ranchingLarge (31-70%)UnknownHigh (continuing)
3 - Energy production & miningSmall (1-10%)Serious - slightHigh (continuing)
4 - Transportation & service corridorsSmall (1-10%)Negligible or <1% pop. declineHigh (continuing)
5 - Biological resource useNegligible (<1%)High (continuing)
5.3 - Logging & wood harvestingHigh (continuing)
6 - Human intrusions & disturbanceSmall (1-10%)Slight or 1-10% pop. declineHigh (continuing)
6.1 - Recreational activitiesHigh (continuing)
6.3 - Work & other activitiesHigh (continuing)
7 - Natural system modificationsNegligible or <1% pop. declineHigh (continuing)
7.1 - Fire & fire suppressionHigh (continuing)
8 - Invasive & other problematic species, genes & diseasesRestricted - smallSerious - slightModerate (short-term)
9 - PollutionLarge - smallUnknownHigh (continuing)
9.3 - Agricultural & forestry effluentsHigh (continuing)
10 - Geological eventsNegligible (<1%)
11 - Climate change & severe weatherPervasive (71-100%)UnknownHigh (continuing)

Roadless Areas (1)
New Mexico (1)
AreaForestAcres
Frisco BoxGila National Forest38,979
References (31)
  1. Adams, R. A. 2003. Bats of the Rocky Mountain West: natural history, ecology, and conservation. University Press of Colorado, Boulder, Colorado. xiii + 289 pp.
  2. American Society of Mammalogists (ASM). 2025. Mammal Diversity Database (Version 1.13) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.10595931. Online. Available: https://www.mammaldiversity.org/
  3. Ammerman, L. K., C. L. Hice, and D. J. Schmidly. 2012. Bats of Texas. Texas A &amp; M University Press, College Station, Texas. xvi + 305 pp.
  4. Arizona Game and Fish Department (AGFD). 2003. <i>Myotis occultus</i>. Unpublished abstract compiled and edited by the Heritage Data Management System, Arizona Game and Fish Department, Phoenix, Arizona. 6 pp.
  5. Arnett, E. B., and E. F. Baerwald. 2013. Impacts of wind energy development on bats: implications for conservation. Pages 435-456 in R. A. Adams and S. C. Pedersen (editors). Bat evolution, ecology, and conservation. Springer Science+Business Media, New York.
  6. Baker, R. J., L. C. Bradley, R. D. Bradley, J. W. Dragoo, M. D. Engstrom, R. S. Hoffman, C. A. Jones, F. Reid, D. W. Rice, and C. Jones. 2003a. Revised checklist of North American mammals north of Mexico, 2003. Museum of Texas Tech University Occasional Papers 229:1-23.
  7. Bradley, R.D., L.K. Ammerman, R.J. Baker, L.C. Bradley, J.A. Cook. R.C. Dowler, C. Jones, D.J. Schmidly, F.B. Stangl Jr., R.A. Van den Bussche and B. Würsig. 2014. Revised checklist of North American mammals north of Mexico, 2014. Museum of Texas Tech University Occasional Papers 327:1-28. Available at: http://www.nsrl.ttu.edu/publications/opapers/ops/OP327.pdf
  8. Calvert, A. W., and S. A. Neiswenter. 2012. Bats in riparian-restoration sites along the lower Colorado River, Arizona. Southwestern Naturalist 57:340-342.
  9. Chung-MacCoubrey, A. 1999 Maternity roosts of bats at the Bosque del Apache National Wildlife Refuge: a preliminary report. USDA Forest Service Proceedings RMRS-P-7.
  10. Findley, J. S. and C. Jones. 1967. Taxonomic relationships of bats of the species <i>Myotis fortidens, M. lucifugus</i> and <i>M. occultus</i>. Journal of Mammalogy 48:429-444.
  11. Fitzgerald, J. P., C. A. Meaney, and D. M. Armstrong. 1994 [1995]. Mammals of Colorado. Denver Museum of Natural History and University Press of Colorado. xiii + 467 pp.
  12. Frey, J. K. 2004. Taxonomy and distribution of the mammals of New Mexico: an annotated checklist. Museum of Texas Tech University Occasional Papers 240. 32 pp.
  13. Geluso, K. 2007. Winter activity of bats over water and along flyways in New Mexico. Southwestern Naturalist 52:482-492.
  14. Geluso, K., and J. N. Mink. 2009. Use of bridges by bats (Mammalia: Chiroptera) in the Rio Grande valley, New Mexico. Southwestern Naturalist 54:421-429.
  15. Hall, E. R. 1981a. The Mammals of North America, second edition. Vols. I &amp; II. John Wiley &amp; Sons, New York, New York. 1181 pp.
  16. Harvey, M. J., J. S. Altenbach, and T. L. Best. 2011. Bats of the United States and Canada. Johns Hopkins University Press, Baltimore, Maryland. 202 pp.
  17. Hayward, B. J. 1963. A maternity colony of <i>Myotis occultus</i>. Journal of Mammalogy 44:279.
  18. Hoffmeister, D. F. 1986. Mammals of Arizona. University of Arizona Press and Arizona Game and Fish Department. 602 pp.
  19. Howell, D. J. 1989. Inventory of known roosts for five species of southwestern bats: <i>Macrotus californicus</i>, <i>Choeronycteris mexicana</i>, <i>Myotis occultus</i>, <i>Myotis auriculus apache</i>, and <i>Idionycteris phyllotis</i>. Unpublished report to U.S. Fish and Wildlife Service, Office of Endangered Species, Albuquerque, New Mexico.
  20. Morgan, C. N., L. K. Ammerman, K. D. Demere, J. B. Doty, Y. J. Nakazawa, and M. R. Mauldin. 2019. Field identification key and guide for bats of the United States of America. Occasional Papers, Museum of Texas Tech University, Number 360. Texas Tech University Natural Science Research Laboratory. 29 pp.
  21. Morrell, T. E., M. J. Rabe, J. C. Devos, Jr., H. Green, and C. R. Miller. 1999. Bats captured in two ponderosa pine habitats in north-central Arizona. Southwestern Naturalist 44:501-506.
  22. Piaggio, A. J., E. W. Valdez, M. A. Bogan, and G. S. Spicer. 2002. Systematics of Myotis occultus (Chiroptera: Vespertilionidae) inferred from sequences of two mitochondrial genes. Journal of Mammalogy 83:386-395.
  23. Rabe, M. J., T. E. Morrell, H. Green, J. C. DeVos, Jr., and C. R. Miller. 1998a. Characteristics of ponderosa pine snag roosts used by reproductive bats in northern Arizona. Journal of Wildlife Management 62:612-621.
  24. Reid, F. A. 2006. A field guide to mammals of North America north of Mexico. Houghton Mifflin Company, Boston.
  25. Schmidly, D. J. 2004. The mammals of Texas. Revised edition. University of Texas Press, Austin. xviii + 501 pp.
  26. Simmons, N. B. and A. L. Cirranello. 2025. Bat Species of the World: A taxonomic and geographic database. Version 1.7. Online. Available: https://batnames.org/
  27. Stager, K. E. 1943b. Remarks on <i>Myotis occultus</i> in California. Journal of Mammalogy 24:197-199.
  28. Valdez, E. W. 2006. Geographic variation in morphology, diet, and ectoparasites of the bat <i>Myotis occultus</i> in New Mexico and southern Colorado. Ph.D. disseration, University of New Mexico, Albuquerque.
  29. Warner, R. M. 1985. Interspecific and temporal dietary variation in an Arizona bat community. Journal of Mammalogy 66:45-51.
  30. Wilson, D. E., and D. M. Reeder (editors). 2005. Mammal species of the world: a taxonomic and geographic reference. Third edition. The Johns Hopkins University Press, Baltimore. Two volumes. 2,142 pp. [As modified by ASM the Mammal Diversity Database (MDD) at https://www.mammaldiversity.org/index.html]
  31. Wilson, D. E., and S. Ruff. 1999. The Smithsonian book of North American mammals. Smithsonian Institution Press, Washington, D.C. 750 pp.