Lasiurus borealis

(Muller, 1776)

Eastern Red Bat

G3Vulnerable (G3G4) Found in 13 roadless areas NatureServe Explorer →
G3VulnerableGlobal Rank
Least concernIUCN
High - mediumThreat Impact
Eastern Red Bat (Lasiurus borealis). Photo by Ed O'Connor, CC BY-NC 4.0, via iNaturalist.
Ed O'Connor, CC BY-NC 4.0
Eastern Red Bat (Lasiurus borealis). Photo by Ed O'Connor, CC BY-NC 4.0, via iNaturalist.
Ed O'Connor, CC BY-NC 4.0
Eastern Red Bat (Lasiurus borealis). Photo by Ed O'Connor, CC BY-NC 4.0, via iNaturalist.
Ed O'Connor, CC BY-NC 4.0
Eastern Red Bat (Lasiurus borealis). Photo by Reuven Martin, CC0 1.0, via iNaturalist.
Reuven Martin, CC0 1.0
Eastern Red Bat (Lasiurus borealis). Photo by Reuven Martin, CC0 1.0, via iNaturalist.
Reuven Martin, CC0 1.0
Eastern Red Bat (Lasiurus borealis). Photo by Sarah Richer, CC BY-NC 4.0, via iNaturalist.
Sarah Richer, CC BY-NC 4.0
Identity
Unique IDELEMENT_GLOBAL.2.799416
Element CodeAMACC05010
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassMammalia
OrderChiroptera
FamilyVespertilionidae
GenusLasiurus
Synonyms
Nycteris borealis
Other Common Names
Chauve-souris rousse (FR) Chauve-souris rousse de l'Est (FR) eastern red bat (EN) Un Murciélago (ES)
Concept Reference
Wilson, D. E., and D. M. Reeder (editors). 2005. Mammal species of the world: a taxonomic and geographic reference. Third edition. The Johns Hopkins University Press, Baltimore. Two volumes. 2,142 pp. [As modified by ASM the Mammal Diversity Database (MDD) at https://www.mammaldiversity.org/index.html]
Taxonomic Comments
New World Lasiurus were placed in the genus Nycteris by Hall (1981), who based the change on nomenclatural (rather than biological) concerns; few if any other authors have followed this change.
Conservation Status
Rank MethodLegacy Rank calculation - Excel v3.1x
Review Date2016-04-04
Change Date2015-03-17
Edition Date2015-03-25
Edition AuthorsHammerson, G.
Threat ImpactHigh - medium
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Rank Reasons
Large range in much of the eastern and central United States and adjacent southern Canada and northeastern Mexico; high availability of roost sites (primarily in trees), but these represent solitary individuals or single females with young; habitat availability has been reduced through historical deforestation, but much habitat remains, and species uses managed forest landscapes and urbanized areas with large trees; population size is probably still large but substantially declining; abundant mortality caused by turbines at wind energy facilities appears to be the major threat, and this threat is expected to greatly increase in the near future; reproductive rate of this species is low, and its ability to sustain the current and anticipated level of wind-energy impact is doubtful.
Range Extent Comments
Range includes the central and eastern United States and adjacent southern Canada (west to British Columbia; Nagorsen and Paterson 2012) and northeastern Mexico, with occasional occurrences on Atlantic/Caribbean islands (Baker et al. 1988, Cryan 2003, Patriquin 2004, Reid 2006). Winter range is smaller than summer range and is mainly in the southeastern United States and northeastern Mexico, with the highest concentrations in coastal Atlantic and Gulf of Mexico regions (Cryan 2003). Individuals in the most northerly portions of the winter range usually are males (Cryan 2003, Mormann and Robbins 2007). Additional records in the southwestern United States extend the distribution farther west that previously recognized (Geluso and Valdez 2019).
Occurrences Comments
The number of distinct occurrences has not been determined using standardized/meaningful criteria, but this species is represented by a widely distributed and large number of collection and observation sites and locations (as defined by IUCN).
Threat Impact Comments
Over the long term, deforestation undoubtedly has reduced the available habitat for this species. However, much suitable forest habitat remains, eastern red bats do not require pristine habitat, and they inhabit urban and semi-urban areas that have ample large hardwood trees.

In British Columbia, "This species continues to be threatened by habitat loss as remaining riparian woodlands are cleared for development. Conservation of riparian areas, which support a diversity of insects, is an important factor." (http://wlapwww.gov.bc.ca/sir/fwh/wld/atlas/species/redbat.html).

This is one of the bat species most commonly killed by turbines at wind energy facilities (Gruver 2002; Johnson et al. 2003; Fiedler 2004; Johnson 2005; Baerwald and Barclay 2009, 2011; Arnett et al. 2008; Cryan 2011; Cryan et al. 2012; Ellison 2012; Jameson and Willis 2012). Arnett and Baerwald (2013) estimated that about 143,000-287,400 eastern red bats were killed at wind energy facilities in the United States and Canada during the period from 2000 to 2011 (22 percent of total bat fatalities). Wind energy is expected to expand from 61,000 MW in 2014 to 350,000 MW by 2030, so the cumulative impact from wind turbines on this species could be devastating. Although the size of the overall eastern red bat population is unknown, the reproductive rate for this species is low, and its ability to sustain the current and anticipated level of impact is doubtful, particularly in light of the additional stressors experienced by this species.

Broadcast application of pesticides to combat forest/tree insect pests potentially has a detrimental impact on this species and its food resources; range-wide population impact is uncertain.

This species is unlikely to be affected by white-nose syndrome (a cold-loving fungus that afflicts bats hibernating in caves and mines).
Ecology & Habitat

Habitat

Habitat includes a wide range of forested and semi-forested areas (e.g. O'Keefe et al. 2009), including developed areas with large trees (e.g., city parks) and some areas subject to intensive forest management (e.g., Elmore et al. 2004). In summer, females tend to occur in warmer lowland regions whereas males are proportionally more common in cooler highlands (Ford et al. 2002). Spring/summer diurnal roosts usually are in foliage of numerous species of large hardwood trees (also pines in some areas; Elmore et al. 2004); suitable sites are open underneath to allow easy exit and entry. Individuals frequently switch roost sites in both summer and winter (Hutchinson and Lacki 2000, Mager and Nelson 2001, Mormann and Robbins 2007). Solitary females roost with young in tree foliage.

In spring and summer in eastern Kentucky, eastern red bats roosted an average of 16.5 meters above ground in the outer foliage of the canopy of 13 species of large hardwood trees, at least 50 meters from forest edges (Hutchinson and Lacki 2000). Most roost trees were on ridge tops of upland forests.

In an intensively managed pine landscape in Mississippi, red bats generally roosted diurnally in foraging areas, which usually contained a reliable water source and open canopy conditions (young open canopy stands, thinned stands, and riparian hardwood stands) (Elmore et al. 2005).

In an urban-rural interface in Indiana, red bats foraged in woodlands and over newly planted tree fields, open water, and pasture lands more than predicted by randomly generated points and avoided highly urban areas such as commercial lands, gravel pits, and transportation corridors (Walters et al. 2007).

In summer in an urban area in Illinois, red bats roosted usually in the foliage or on the bark of large deciduous trees, sometimes in leaf litter, dense grass, or shingles of houses (Mager and Nelson 2001).

In southwestern Missouri, winter roosts were in eastern red-cedar or hardwoods, on the south side of trees, on south-facing slopes; bats switched from tree roosts to leaf litter roosts when ambient temperatures approached or fell below freezing (Mormann and Robbins 2007). Similar behavior has been recorded elsewhere (e.g., Rodrigue et al. 2001).

Ecology

Basically solitary.

Reproduction

Copulates August-October in North America (also in spring in some areas?). Gestation lasts 60-70 (also reported as 80-90) days, after delayed fertilization. Litter of 1-5 (average 2), born late May to mid-June (or July) in North America. Young can fly at 3-4 weeks, weaned at 5-6 weeks. Sexually mature during second month.
Terrestrial Habitats
Forest - HardwoodForest - MixedWoodland - HardwoodWoodland - MixedSuburban/orchardAerial
Palustrine Habitats
RiparianAerial
Other Nations (2)
CanadaN4B,NUM
ProvinceRankNative
ManitobaS3BYes
Nova ScotiaSUB,S1MYes
SaskatchewanS4BYes
OntarioS3Yes
AlbertaS3B,SNRMYes
British ColumbiaSUYes
New BrunswickSUB,S2MYes
QuebecS1BYes
United StatesN5
ProvinceRankNative
MaineSUYes
WisconsinS3Yes
ConnecticutS3BYes
TexasS4Yes
MassachusettsS3Yes
Rhode IslandSNRYes
MissouriS4Yes
GeorgiaS5Yes
New HampshireS3BYes
OklahomaSNRYes
New JerseyS3Yes
West VirginiaS3Yes
OhioSNRYes
North CarolinaS5Yes
New MexicoS3NYes
PennsylvaniaS4Yes
ColoradoS2BYes
VirginiaS4Yes
IllinoisS5Yes
ArizonaSNRYes
South DakotaS5Yes
KentuckyS5Yes
ArkansasS5Yes
MontanaS3BYes
MinnesotaSNRYes
MississippiS4Yes
KansasS5BYes
DelawareS5Yes
IndianaS3Yes
New YorkS3BYes
NebraskaS3Yes
District of ColumbiaS4Yes
North DakotaSNRYes
FloridaSNRYes
South CarolinaS3Yes
MarylandS3Yes
AlabamaS5Yes
TennesseeS5Yes
LouisianaS4Yes
VermontS4BYes
IowaS4Yes
WyomingS3BYes
MichiganS5Yes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentNegligible (<1%)High (continuing)
2 - Agriculture & aquacultureNegligible (<1%)High (continuing)
3 - Energy production & miningLarge - restrictedSerious or 31-70% pop. declineHigh (continuing)
3.3 - Renewable energyHigh (continuing)
4 - Transportation & service corridorsSmall (1-10%)Negligible or <1% pop. declineHigh (continuing)
5 - Biological resource useSmall (1-10%)Serious - slightHigh (continuing)
5.3 - Logging & wood harvesting
6 - Human intrusions & disturbanceSmall (1-10%)Negligible or <1% pop. declineHigh (continuing)
7 - Natural system modificationsNegligible (<1%)High (continuing)
8 - Invasive & other problematic species, genes & diseasesLarge (31-70%)Negligible or <1% pop. declineHigh (continuing)
9 - PollutionRestricted - smallUnknownHigh (continuing)
10 - Geological eventsNegligible (<1%)
11 - Climate change & severe weatherPervasive (71-100%)UnknownHigh (continuing)

Roadless Areas (13)
Arkansas (2)
AreaForestAcres
Dismal CreekOzark-St. Francis National Forest9,160
Richland CreekOzark-St. Francis National Forest571
Georgia (2)
AreaForestAcres
Kelly RidgeChattahoochee National Forest8,325
Miller CreekChattahoochee National Forest701
North Carolina (1)
AreaForestAcres
Laurel MountainPisgah National Forest5,683
Tennessee (3)
AreaForestAcres
Bald MountainCherokee National Forest11,743
Brushy RidgeCherokee National Forest7,469
Sycamore CreekCherokee National Forest6,984
Virginia (3)
AreaForestAcres
Beards MountainGeorge Washington National Forest7,505
Brush MountainJefferson National Forest6,002
Kelley MountainGeorge Washington National Forest7,590
West Virginia (2)
AreaForestAcres
Gauley MountainMonongahela National Forest13,285
Middle MountainMonongahela National Forest19,020
References (86)
  1. Adams, R. A. 2003. Bats of the Rocky Mountain West: natural history, ecology, and conservation. University Press of Colorado, Boulder, Colorado. xiii + 289 pp.
  2. American Society of Mammalogists (ASM). 2025. Mammal Diversity Database (Version 1.13) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.10595931. Online. Available: https://www.mammaldiversity.org/
  3. Anonymous. 2002. Plan for biodiversity management and sustainable development of the Turks and Caicos RAMSAR site. Turks & Caicos National Trust, UK Overseas Territories Conservation Forum, and CABI Bioscience, http://www.ukotcf.org/pdf/TCIRamMP/Contents%20and%20Summary.pdf.
  4. Arnett, E. B., and E. F. Baerwald. 2013. Impacts of wind energy development on bats: implications for conservation. Pages 435-456 in R. A. Adams and S. C. Pedersen (editors). Bat evolution, ecology, and conservation. Springer Science+Business Media, New York.
  5. Arnett, E. B., W. K. Brown, W. P. Erickson, J. K. Fiedler, B. L. Hamilton, T. H. Henry, A. Jain, G. D. Johnson, J. Kerns, R. R. Koford, C. P. Nicholson, T. J. Connell, M. D. Piorkowski and R. D. Tankersley. 2008. Patterns of bat fatalities at wind energy facilities in North America. Journal of Wildlife Management 72(1): 61-78.
  6. Baerwald, E. F., and R.M.R. Barclay. 2009. Geographic variation in activity and fatality of migratory bats at wind energy facilities. Journal of Mammalogy 90:1341-1349.
  7. Baerwald, E. F., and R.M.R. Barclay. 2011. Patterns of activity and fatality of migratory bats at a wind energy facility in Alberta, Canada. Journal of Wildlife Management 75:11013-1114.
  8. Baker, R. H. 1983. Michigan mammals. Michigan State University Press. 642 pp.
  9. Baker, R. J., J. C. Patton, H. H. Genoways, and J. W. Bickham. 1988. Genic studies of Lasiurus (Chiroptera: Vespertilionidae). Occas. Pap. Mus. Texas Tech. Univ. 117:1-15.
  10. Baker, R. J., L. C. Bradley, R. D. Bradley, J. W. Dragoo, M. D. Engstrom, R. S. Hoffman, C. A. Jones, F. Reid, D. W. Rice, and C. Jones. 2003a. Revised checklist of North American mammals north of Mexico, 2003. Museum of Texas Tech University Occasional Papers 229:1-23.
  11. Banfield, A. W. F. 1974. The mammals of Canada. University of Toronto Press, Toronto, Canada. 438 pp.
  12. Barbour, R. W., and W. H. Davis. 1969. Bats of America. The University of Kentucky Press, Lexington, Kentucky. 286 pp.
  13. Bárquez, R. M., M. A. Mares, and J. K. Braun. 1999. The bats of Argentina. Special Publications, Museum of Texas Tech University 42:1-275.
  14. Bradley, R.D., L.K. Ammerman, R.J. Baker, L.C. Bradley, J.A. Cook. R.C. Dowler, C. Jones, D.J. Schmidly, F.B. Stangl Jr., R.A. Van den Bussche and B. Würsig. 2014. Revised checklist of North American mammals north of Mexico, 2014. Museum of Texas Tech University Occasional Papers 327:1-28. Available at: http://www.nsrl.ttu.edu/publications/opapers/ops/OP327.pdf
  15. Caire, W., J. D. Tyler, B. P. Glass, and M. A. Mares. 1989. Mammals of Oklahoma. University of Oklahoma Press, Norman. Oklahoma. 567 pp.
  16. Carter, T. C. 1998. The foraging ecology of three species of bats at the Savannah River Site, South Carolina. M.S. thesis, University of Georgia, Athens. 74 pp.
  17. Clawson, R. L., R. K. LaVal, M. L. LaVal, and W. Claire. 1980. Clustering behavior of hibernating <i>Myotis sodalis</i> in Missouri, USA. Journal of Mammalogy 61:245-53.
  18. Cryan, P. M. 2003. Seasonal distribution of migratory tree bats (<i>Lasiurus </i>and <i>Lasionycteris</i>) in North America. Journal of Mammalogy 84:579-593.
  19. Cryan, P. M. 2011. Wind turbines as landscape impediments to the migratory connectivity of bats. Environmental Law 41(2):355-370.
  20. Cryan, P. M., J. W. Jameson, E. F. Baerwald, C.K.R. Willis, R.M.R. Barclay, E. A. Snider, and E. G. Crichton. 2012. Evidence of late-summer mating readiness and early sexual maturation in migratory tree-roosting bats found dead at wind turbines. PLoS ONE 7(10): e47586. doi:10.1371/journal.pone.0047586.
  21. Elliott, W. R. 2003. A guide to Missouri's cave life: 70 species brought to life. Missouri Department of Conservation. 37 pp.
  22. Ellison L. E. 2012. Bats and wind energy: a literature synthesis and annotated bibliography. U.S. Geological Survey. Open-File Report No. 2012-1110.
  23. Elmore, L. W., D. A. Miller, and F. J. Vilella. 2004. Selection of diurnal roosts by red bats (<i>Lasiurus borealis</i>) in an intensively managed pine forest in Mississippi. Forest Ecology and Management 199:11-20.
  24. Elmore, L. W., D. A. Miller, and F. J. Vilella. 2005. Foraging area size and habitat use by red bats (Lasiurus borealis) in an intensively managed pine landscape in Mississippi. American Midland Naturalist 153:405-417.
  25. Fellers, G. M., and E. D. Pierson. 2002. Habitat use and foraging behavior of Townsend's Big-eared Bat (<i>Corynorhinus townsendii</i>) in coastal California. Journal of Mammalogy 83:167-177.
  26. Fiedler, J. K. 2004, Assessment of bat mortality and activity at Buffalo Mountain Windfarm, eastern Tennessee, Master's thesis, University of Tennessee, Knowville. 166 pp.
  27. Flinn, J. 2009. Winter roosting behavior of red bats (<i>Lasiurus borealis</i>): habitat use, microclimate, and effects of ambient temperature on roost choice. M.S. thesis. Missouri State University, Springfield, MO.
  28. Ford, W. M., M. A. Menzel, J. M. Menzel, and D. J. Welch. 2002. Influence of summer temperature on sex rations in eastern red bats (Lasiurus borealis). American Midland Naturalist 147:179-184.
  29. Furlonger, C. L., H. J. Dewar, and M. B. Fenton. 1987. Habitat use by foraging insectivorous bats. Canadian J. Zool. 65:284-288.
  30. Garner, J. D., and J. E. Gardner. 1992. Determination of summer distribution and habitat utilization of the Indiana bat (<i>Myotis sodalis</i>) in Illinois. Illinois Department of Conservation. Final Report, Project E-3. Springfield, IL, 23 pp.
  31. Geluso, K., and E. W. Valdez. 2019. First Records of the Eastern Red Bat (<i>Lasiurus borealis</i>) in Arizona, Utah, and Western New Mexico. Occasional Papers, Museum of Texas Tech University 361:1-16.
  32. Godin, A. J. 1977. Wild mammals of New England. Johns Hopkins University Press, Baltimore. 304 pp.
  33. Griffin, D. R. 1940b. Migrations of New England bats. Bulletin of the Museum of Comparative Zoology 86:217-246.
  34. Gruver, J. C. 2002. Assessment of bat community structure and roosting habitat preferences for the hoary bat (<i>Lasiurus cinereus</i>) near Foote Creek Rim, Wyoming. Master's thesis, University of Wyoming, Laramie. 164 pp.
  35. Hall, E. R. 1981a. The Mammals of North America, second edition. Vols. I &amp; II. John Wiley &amp; Sons, New York, New York. 1181 pp.
  36. Hamilton, W. J., Jr., and J. O. Whitaker, Jr. 1979. Mammals of the eastern United States. Cornell Univ. Press, Ithaca, New York. 346 pp.
  37. Hart, J. A., G. L. Kirkland, Jr., and S. C. Grossman. 1993. Relative abundance and habitat use by tree bats, LASIURUS spp., in southcentral Pennsylvania. Can. Field-Nat. 107:208-212.
  38. Hatch, S. K., E. E. Connelly, T. J. Divoll, I. J. Stenhouse, and K. A. Williams. 2013. Offshore observations of eastern red bats (<i>Lasiurus borealis</i>) in the mid-Atlantic United States using multiple survey methods. PLoS ONE 8(12): e83803. doi:10.1371/journal.pone.0083803.
  39. Hebda, A.J. 2011. List of mammals of Nova Scotia (including synonyms used in the literature relating to Nova Scotia) (revision 2) 24 July 2011. Nova Scotia Museum Collections Unit, Halifax, Nova Scotia. 24 pp. Online. Available: https://naturalhistory.novascotia.ca/sites/default/files/inline/images/names_and_synonyms_ver3.pdf
  40. Hickey, M. B. C., and M. B. Fenton. 1990. Foraging by red bats (LASIURUS BOREALIS): do intraspecific chases mean territoriality? Can. J. Zool. 68:2477-2482.
  41. Hutchinson, J. T., and M. J. Lacki. 1999. Foraging behavior and habitat use of red bats in mixed mesophytic forests of the Cumberland Plateau, Kentucky. Pages 171-177 in J. W. Stringer and D. L. Loftis, editors. 12th Central Hardwood Forest Conference, U.S. Forest Service, Southern Experiment Station, Asheville, North Carolina.
  42. Hutchinson, J. T., and M. J. Lacki. 2000. Selection of day roosts by red bats in mixed mesophytic forests. Journal of Wildlife Management 64:87-94.
  43. Jackson, H. H. 1961. Mammals of Wisconsin. University of Wisconsin Press, Madison. 504 pp.
  44. Jameson, J.W. and C.K. Willis. 2012. Bat mortality at a wind power facility in central Canada. Northwestern Naturalist 93(3): 194-202.
  45. Johnson, G. D. 2005. A review of bat mortality at wind-energy developments in the United States. Bat Research News 46:45-49.
  46. Johnson, G. D., W. P. Erickson, M. D. Strickland, M. F. Shepherd, and S. A. Sarappo. 2003. Mortality of bats at a large-scale wind power development at Buffalo Ridge, Minnesota. American Midland Naturalist 150:332-342.
  47. Jones, J. K., Jr., R. S. Hoffman, D. W. Rice, C. Jones, R. J. Baker, and M. D. Engstrom. 1992a. Revised checklist of North American mammals north of Mexico, 1991. Occasional Papers, The Museum, Texas Tech University, 146:1-23.
  48. Kiser, J. D. and C. L. Elliott. 1996. Foraging habitat, food habits, and roost tree characteristics of the Indiana bat (<i>Myotis sodalis</i>) during autumn in Johnson County, Kentucky. Final report, Kentucky Department of Fish and Wildlife Resources, Frankfort, Kentucky. 65pp.
  49. Koopman, K. F., and G. F. McCracken. 1998. The taxonomic status of <i>Lasiurus</i> (Chiroptera: Vespertilionidae) in the Galapagos Islands. American Museum Novitates 3243:1-6.
  50. LaVal, R.K. and M.L. LaVal. 1979. Notes on reproduction, behavior, and abundance of the red bat, LASIURUS BOREALIS. J. Mammalogy, 60(1):209-212.
  51. Linzey, D. W. 1998. The mammals of Virginia. McDonald and Woodward, Blacksburg, VA. 459 pp.
  52. Linzey, D.W. 2016. Mammals of Great Smoky Mountains National Park: 2016 revision. Southeastern Naturalist 15(Monograph 8):1–93.
  53. Mager, K. J., and T. A. Nelson. 2001. Roost-site selection by eastern red bats (Lasiurus borealis). American Midland Naturalist 145:120-126.
  54. McClure, H.E. 1942. Summer activities of bats (genus Lasiurus) in Iowa. Jour. Mamm. 23(4):430-434.
  55. Menzel, M. A., R. Odom, S. Owen, W. M. Ford, B. R. Chapman, K. V. Miller, J. Edwards, and P. Wood. 1999b. Investigation of foraging habitat use by bats with a focus on Northern Long-eared Myotis (<i>Myotis septentrionalis</i>): a comparison of methods. IN M. K. Clark, editor. Abstracts from the 1999 Colloquium on the conservation of mammals in the Southeastern United States. Available at: http://www.batworkinggroups.org/sbdnnews.htm. Accessed 2001-06-12.
  56. Milam-Dunbar, M. B. 2005. Ecophysiology of hibernating eastern red bats (<i>Lasiurus borealis</i>). Ph.D. dissertation, Missouri State University.
  57. Morales, J. C., and J. W. Bickham. 1995. Molecular systematics of the genus <i>Lasiurus </i>(Chiroptera: Vespertilionidae) based on restriction-site maps of the mitochondrial ribosomal genes. Journal of Mammalogy 76:730-749.
  58. Morgan, C. N., L. K. Ammerman, K. D. Demere, J. B. Doty, Y. J. Nakazawa, and M. R. Mauldin. 2019. Field identification key and guide for bats of the United States of America. Occasional Papers, Museum of Texas Tech University, Number 360. Texas Tech University Natural Science Research Laboratory. 29 pp.
  59. Mormann, B.M. and L.W. Robbins. 2007. Winter roosting ecology of eastern red bats in southwest Missouri. The Journal of Wildlife Management 71(1):213-217.
  60. Nagorsen, D. W., and B. Paterson. 2012. An update on the status of red bats, <i>Lasiurus blossevilli</i>i and <i>Lasiurus borealis</i>, in British Columbia. Northwestern Naturalist 93:235-237.
  61. O'Keefe, J. M., S. C. Loeb, J. D. Lanham, and H. S. Hill, Jr. 2009. Macrohabitat factors affect day roost selection by eastern red bats and eastern pipistrelles in the southern Appalachian Mountains, USA. Forest Ecology and Management 257:1757-1763.
  62. Oliver, G. V. 2000. The bats of Utah: a literature review. Publication No. 00-14, Utah Division of Wildlife Resources, Salt Lake City.
  63. Padgett, T. M., and R. K. Rose. 1991. Bats (Chiroptera: Vespertilionidae) of the Great Dismal Swamp of Virginia and North Carolina. Brimleyana 17:17-25.
  64. Patriquin, K. J. 2004. Red bat (<i>Lasiurus borealis</i>) captured in northeastern Alberta. Northwestern Naturalist 85:28-30.
  65. Perry, R. W. 2013. Potential energy expenditure by litter-roosting bats associated with temperature under leaf litter during winter. Journal of Thermal Biology 38:467-473.
  66. Ransome, R. 1990. The natural history of hibernating bats. Christopher Helm, London. xxi + 235 pp.
  67. Reid, F. A. 1997. A field guide to the mammals of Central America and southeast Mexico. Oxford University Press, Incorporated New York, New York. 334 pp.
  68. Reid, F. A. 2006. A field guide to mammals of North America north of Mexico. Houghton Mifflin Company, Boston.
  69. Rodrigue, J.L,, T. M. Schuler, and M. A. Menzel. 2001. Observations of bat activity during prescribed burning in West Virginia. Bat Research News 42:48-49.
  70. Rodríguez-Durán, A. and T.H. Kunz. 2001. Biogeography of West Indian bats: An ecological perspective. Pp. 355-368 In C. A. Woods and F. E. Sergile (eds.), Biogeography of the West Indies. CRC Press, Boca Raton, FL, USA.
  71. Schmidly, D. J. 1991. The bats of Texas. Texas A & M University Press, College Station, Texas. 188 pp.
  72. Schwartz, C. W., and E. R. Schwartz. 1981. The wild mammals of Missouri. University of Missouri Press, Columbia. 356 pp.
  73. Segers, J. L., A. E. Irwin, L. J. Farrow, L.N.L. Johnson, and H. G. Broders. 2013. First records of <i>Lasiurus cinereus</i> and <i>L. borealis</i> (Chiroptera: Vespertilionidae) on Cape Breton Island, Nova Scotia, Canada. Northeastern Naturalist 20:N14-N15.
  74. Shump, K. A., Jr., and A. U. Shump. 1982b. LASIURUS BOREALIS. Am. Soc. Mamm., Mammalian Species No. 183:1-6.
  75. Simmons, N. B. and A. L. Cirranello. 2025. Bat Species of the World: A taxonomic and geographic database. Version 1.7. Online. Available: https://batnames.org/
  76. Smallwood, K. S. 2013. Comparing bird and bat fatality-rate estimates among North American wind-energy projects. Wildlife Society Bulletin 37:19-33.
  77. Tirira, D. 1999. Mamíferos del Ecuador. Museo de Zoología, Pontificia Universidad Católica del Ecuador, Quito.
  78. Tuttle, M. D. 1976a. Population ecology of the gray bat (<i>Myotis grisescens</i>): philopatry, timing, and patterns of movement, weight loss during migration, and seasonal adaptive strategies. University of Kansas Museum of Natural History Occasional Papers (54):1-38.
  79. U.S. Fish and Wildlife Service (USFWS). 1999. Agency draft Indiana Bat (<i>Myotis sodalis</i>) revised recovery plan. U.S. Fish and Wildlife Service, Fort Snelling, Minnesota. 53 pp.
  80. van Zyll de Jong, C.G. 1985. Handbook of Canadian Mammals. Vol. II, Bats. National Museum of Natural Sciences, National Museums of Canada, Ottawa, Canada. 212 pp.
  81. Walters, B. L., C. M. Ritzi, D. W. Sparks, and J. O. Whitaker, Jr. 2007. Foraging behavior of eastern red bats (<i>Lasiurus borealis</i>) at an urban-rural interface. American Midland Naturalist 157:365-373.
  82. Whitaker, J. O., Jr., R. K. Rose, and T. M. Padgett. 1997. Food of the red bat <i>Lasiurus borealis</i> in winter in the Great Dismal Swamp, North Carolina and Virginia. American Midland Naturalist 137:408-411.
  83. Wilson, D. E., and D. M. Reeder (editors). 1993. Mammal species of the world: a taxonomic and geographic reference. Second edition. Smithsonian Institution Press, Washington, DC. xviii + 1206 pp. Available online at: http://www.nmnh.si.edu/msw/.
  84. Wilson, D. E., and D. M. Reeder (editors). 2005. Mammal species of the world: a taxonomic and geographic reference. Third edition. The Johns Hopkins University Press, Baltimore. Two volumes. 2,142 pp. [As modified by ASM the Mammal Diversity Database (MDD) at https://www.mammaldiversity.org/index.html]
  85. Wilson, D. E., and S. Ruff. 1999. The Smithsonian book of North American mammals. Smithsonian Institution Press, Washington, D.C. 750 pp.
  86. Winhold, L., A. Kurta, and R. Foster. 2008. Long-term change in an assemblage of North American bats: are eastern red bats declining? Acta Chiropterologica 10:359-366.