Agkistrodon contortrix

(Linnaeus, 1766)

Eastern Copperhead

G5Secure Found in 87 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Least concernIUCN
Medium - lowThreat Impact
Eastern Copperhead (Agkistrodon contortrix). Photo by Brent Cox, CC BY 4.0, via iNaturalist.
Brent Cox, CC BY 4.0
Eastern Copperhead (Agkistrodon contortrix). Photo by Jennifer Schultz, CC0 1.0, via iNaturalist.
Jennifer Schultz, CC0 1.0
Eastern Copperhead (Agkistrodon contortrix). Photo by acartes, CC BY 4.0, via iNaturalist.
acartes, CC BY 4.0
Identity
Unique IDELEMENT_GLOBAL.2.960824
Element CodeARADE01040
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassReptilia
OrderSquamata
FamilyViperidae
GenusAgkistrodon
Other Common Names
copperhead (EN)
Concept Reference
Burbrink, F. T., and T. J. Guiher. 2014 [2015]. Considering gene flow when using coalescent methods to delimit lineages of North American pitvipers of the genus Agkistrodon. Zoological Journal of the Linnean Society 173(2):505-526.
Taxonomic Comments
Using multi-locus nuclear data, Burbrink and Guiher (2015) partially confirmed previous mitochondrial hypotheses (Guiher and Burbrink 2008) that the two North American species (A. contortrix and A. piscivorus) each consist of multiple species-level taxa. Subspecies are not recognized; this species comprises the previously recognized subspecies Agkistrodon contortrix contortrix, Agkistrodon contortrix mokasen, and Agkistrodon contortrix phaeogaster (in part).
Conservation Status
Rank Method Rank calculation - Biotics v2
Review Date2025-10-07
Change Date1996-10-31
Edition Date2025-10-07
Edition AuthorsHammerson, G. (2006); rev. R. L. Gundy (2025)
Threat ImpactMedium - low
Range Extent200,000-2,500,000 square km (about 80,000-1,000,000 square miles)
Number of Occurrences> 300
Rank Reasons
This species has a wide distribution across the eastern U.S. The population is abundant and the species is somewhat tolerant of disturbance and fragmentation.
Range Extent Comments
The geographic range extends from southern New England to northern Florida, and west through the southern Great Lakes states and southern Iowa to southeastern Nebraska, eastern Kansas, eastern Oklahoma, and eastern Texas (Burbrink and Guiher 2015, Powell et al. 2016). Elevational range extends from near sea level to above 1,500 meters. Using Global Biodiversity Information Facility (GBIF) (2025) records, range extent is estimated to be 2.4 million km² (RARECAT 2025).
Occurrences Comments
This species is represented by a very large number of occurrences (subpopulations) (Campbell and Lamar 2004).
Threat Impact Comments
Habitat loss due to development and agriculture is an ongoing threat. This species can be quite tolerant to fragmentation and human disturbance (Carrasco-Harris et al. 2020), although habitat fragmentation can reduce overall abundance.
Ecology & Habitat

Habitat

Copperheads are often in or near deciduous forest in hilly situations, usually in the vicinity of rock outcrops; they occur also on floodplains and at edges of swamps in the south and in mesic situations near water in the arid west. Hibernation generally occurs in dens among rocks, or in caves, animal burrows, under objects, in hollow logs or stumps, or in similar sites. Usually copperheads area in areas with abundant surface cover such as rocks, logs, stumps, or leaf litter. They are mainly terrestrial but sometimes climb into vegetation up to a few meters above ground. In the east at least, gravid females select rocky areas that are more open and have warmer soil temperatures than those used by nongravid individuals (Reinert, cited by Ernst 1992).

Ecology

Hibernates communally (especially in north) or singly. In Kansas, population density in fall was estimated at about 13/ha, with perhaps 2-4 times this many under optimal conditions; home range size was about 10 ha for males and 3.4 ha for females (Fitch 1960).

Reproduction

Most copulations occur in spring and late summer-early fall. Births occur mainly in August or September in most areas, but may occur as early as July or as late as November in some areas. In Kansas, individual females evidently give birth usually in alternate years, July-early October; males sexually mature in 2nd summer, most females in 3 years (Fitch 1960). In southern Texas, individual females may produce young in consecutive years (Vermersch and Kuntz, 1986, Snakes of south central Texas, Eakin Press, Austin). Litter size up to 21 (most often 4-8) in east and north, average of 5.3 in Kansas, usually not more than 3-4 in Trans-Pecos subspecies PICTIGASTER.
Terrestrial Habitats
Forest - HardwoodForest - MixedWoodland - HardwoodWoodland - MixedSavannaOld fieldDesertBare rock/talus/screeCliff
Palustrine Habitats
Riparian
Other Nations (1)
United StatesN5
ProvinceRankNative
IowaS1Yes
FloridaS2Yes
NebraskaS2Yes
MississippiS5Yes
North CarolinaS5Yes
VirginiaS5Yes
West VirginiaS3Yes
TexasS5Yes
GeorgiaS5Yes
ConnecticutS3Yes
MissouriS5Yes
AlabamaS5Yes
MassachusettsS1Yes
South CarolinaS5Yes
New JerseyS3Yes
OklahomaSNRYes
DelawareS1Yes
LouisianaS5Yes
New YorkS3Yes
KentuckyS5Yes
MarylandS5Yes
TennesseeS5Yes
OhioSNRYes
District of ColumbiaS1Yes
PennsylvaniaS3Yes
ArkansasSNRYes
IndianaS4Yes
IllinoisS4Yes
KansasS5Yes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentLarge - restrictedModerate - slightHigh (continuing)
1.1 - Housing & urban areasLarge - restrictedModerate - slightHigh (continuing)
1.2 - Commercial & industrial areasLarge - restrictedModerate - slightHigh (continuing)
2 - Agriculture & aquacultureRestricted (11-30%)Moderate - slightHigh (continuing)
2.1 - Annual & perennial non-timber cropsRestricted (11-30%)Moderate - slightHigh (continuing)

Roadless Areas (87)
Alabama (2)
AreaForestAcres
Blue MountainTalladega National Forest4,986
Oakey MountainTalladega National Forest6,129
Arkansas (11)
AreaForestAcres
Bear MountainOuachita National Forest1,910
Blue MountainOuachita National Forest9,755
Brush HeapOuachita National Forest4,205
Devils CanyonOzark-St. Francis National Forest1,877
Dismal CreekOzark-St. Francis National Forest9,160
East ForkOzark-St. Francis National Forest13,037
Gee CreekOzark-St. Francis National Forest7,957
Little BlakelyOuachita National Forest3,342
Pedestal RocksOzark-St. Francis National Forest21,957
PenhookOzark-St. Francis National Forest6,566
Richland CreekOzark-St. Francis National Forest571
Georgia (12)
AreaForestAcres
Boggs CreekChattahoochee National Forest2,073
Cedar MountainChattahoochee National Forest1,083
Ellicott Rock AdditionChattahoochee National Forest690
Helton CreekChattahoochee National Forest2,348
Joe GapChattahoochee National Forest5,321
Ken MountainChattahoochee National Forest527
Lance CreekChattahoochee National Forest9,025
Miller CreekChattahoochee National Forest701
Pink KnobChattahoochee National Forest12,127
Rocky MountainChattahoochee National Forest4,269
Sarah's CreekChattahoochee National Forest6,888
Turner CreekChattahoochee National Forest1,495
Illinois (1)
AreaForestAcres
Burke BranchShawnee National Forest6,231
Indiana (1)
AreaForestAcres
Mogan RidgeHoosier National Forest8,435
Kentucky (1)
AreaForestAcres
WolfpenDaniel Boone National Forest2,835
Louisiana (1)
AreaForestAcres
Saline Bayou W & S River CorridorKisatchie National Forest5,355
Missouri (2)
AreaForestAcres
Big Creek Rare II Study AreaMark Twain National Forest9,277
Swan Creek Rare II Study AreaMark Twain National Forest7,310
North Carolina (15)
AreaForestAcres
Bald MountainPisgah National Forest11,085
Big Indian (addition)Nantahala National Forest1,155
Catfish Lake NorthCroatan National Forest11,299
Catfish Lake South - ACroatan National Forest217
Harper CreekPisgah National Forest7,325
Jarrett CreekPisgah National Forest7,485
Laurel MountainPisgah National Forest5,683
Linville Gorge AdditionPisgah National Forest2,809
Lost CovePisgah National Forest5,944
Overflow CreekNantahala National Forest3,379
Pond Pine BCroatan National Forest2,961
Sheep Ridge AdditionCroatan National Forest5,808
South Mills RiverPisgah National Forest8,588
Tusquitee BaldNantahala National Forest13,670
Wilson CreekPisgah National Forest4,863
South Carolina (2)
AreaForestAcres
Ellicott Rock 2Sumter National Forest517
Hellhole ExtFrancis Marion National Forest891
Tennessee (6)
AreaForestAcres
Bald MountainCherokee National Forest11,743
Beaver Dam CreekCherokee National Forest5,070
Brushy RidgeCherokee National Forest7,469
Flint Mill GapCherokee National Forest9,494
Sampson Mountain AdditionCherokee National Forest3,064
Stone MountainCherokee National Forest5,367
Texas (3)
AreaForestAcres
Big WoodsNational Forests in Texas1,320
Little Lake CreekNational Forests in Texas596
Winters BayouNational Forests in Texas730
Virginia (28)
AreaForestAcres
Adams PeakGeorge Washington National Forest7,135
Barbours Creek AdditionJefferson National Forest733
Beards MountainGeorge Washington National Forest7,505
Broad RunJefferson National Forest10,971
Brush MountainJefferson National Forest6,002
Elliott KnobGeorge Washington National Forest9,380
Gum RunGeorge Washington National Forest12,620
Hoop HoleJefferson National Forest4,652
James River AdditionJefferson National Forest1,140
JerkemtightGeorge Washington National Forest16,687
Kelley MountainGeorge Washington National Forest7,590
Little RiverGeorge Washington National Forest27,292
Little Walker MountainJefferson National Forest9,818
Mill MountainGeorge Washington National Forest10,840
Mt. PleasantGeorge Washington National Forest8,933
North MountainJefferson National Forest8,377
Northern MassanuttenGeorge Washington National Forest9,444
Oak KnobGeorge Washington National Forest10,882
Oliver MountainGeorge Washington National Forest13,090
Patterson MountainJefferson National Forest4,865
Peters Mountain Addition A (VA)Jefferson National Forest1,268
Ramseys Draft AdditionGeorge Washington National Forest12,781
Seng MountainJefferson National Forest6,428
Shawvers Run AdditionJefferson National Forest1,927
SkidmoreGeorge Washington National Forest5,641
Southern MassanuttenGeorge Washington National Forest11,985
The PriestGeorge Washington National Forest5,737
Three RidgesGeorge Washington National Forest4,745
West Virginia (2)
AreaForestAcres
Dry River (WV)George Washington National Forest7,331
Peters Mountain Addition A (WV)Jefferson National Forest343
References (44)
  1. Ashton, R. E., Jr., and P. S. Ashton. 1981. Handbook of reptiles and amphibians of Florida. Part One: The Snakes. Windward Publishing Company, Miami, Florida. 176 pp.
  2. Barbour, R. W. 1971. Amphibians and reptiles of Kentucky. Univ. Press of Kentucky, Lexington. x + 334 pp.
  3. Behler, J. L., and F. W. King. 1979. The Audubon Society field guide to North American reptiles and amphibians. Alfred A. Knopf, New York. 719 pp.
  4. Burbrink, F. T., and T. J. Guiher. 2014 [2015]. Considering gene flow when using coalescent methods to delimit lineages of North American pitvipers of the genus <i>Agkistrodon</i>. Zoological Journal of the Linnean Society 173(2):505-526.
  5. Campbell, J. A., and D. H. Whitmore, Jr. 1989. A comparison of the skin keratin biochemistry in vipers with comments on its systematic value. Herpetologica 45:242-249.
  6. Campbell, J. A., and E. D. Brodie, Jr., editors. 1992. Biology of the pit vipers. Selva, Tyler, Texas.
  7. Campbell, J. A., and W. W. Lamar. 1989. The venomous reptiles of Latin America. Comstock Publ. Associates, Division of Cornell Univ. Press, Ithaca, New York. xii + 425 pp.
  8. Campbell, J. A., and W. W. Lamar. 2004. The venomous reptiles of the Western Hemisphere. 2 volumes. Cornell University Press.
  9. Carrasco-Harris, M. F., D. Bowman, S. Reichling, and J. A. Cole. 2020. Spatial ecology of copperhead snakes (<i>Agkistrodon contortrix</i>) in response to urban park trails. Journal of Urban Ecology 6(1): 1–7. doi: 10.1093/jue/juaa007
  10. Chiasson, R. B., D. L. Bentley, and C. H. Lowe. 1989. Scale morphology in <i>Agkistrodon</i>, and closely related crotaline genera. Herpetologica 45:430-438.
  11. Collins, J. T. 1982. Amphibians and reptiles in Kansas. Second edition. Univ. Kansas Mus. Nat. Hist., Pub. Ed. Ser. 8. xiii + 356 pp.
  12. Collins, J. T. 1990. Standard common and current scientific names for North American amphibians and reptiles. 3rd ed. Society for the Study of Amphibians and Reptiles. Herpetological Circular No. 19. 41 pp.
  13. Collins, J. T. 1993. Amphibians and reptiles in Kansas. Third edition, revised. University of Kansas Museum of Natural History, Public Education Series No. 13. xx + 397 pp.
  14. Conant, R. and J. T. Collins. 1991. A field guide to reptiles and amphibians: eastern and central North America. Third edition. Houghton Mifflin Co., Boston, Massachusetts. 450 pp.
  15. Conant, R., and J. T. Collins. 1998. A field guide to reptiles and amphibians: eastern and central North America. Third edition, expanded. Houghton Mifflin Co., Boston, Massachusetts. 616 pp.
  16. Crother, B. I. (editor). 2008. Scientific and standard English names of amphibians and reptiles of North America north of Mexico, with comments regarding confidence in our understanding. Sixth edition. Society for the Study of Amphibians and Reptiles Herpetological Circular 37:1-84. Online with updates at: http://www.ssarherps.org/pages/comm_names/Index.php
  17. Crother, B. I. (editor). 2012. Scientific and standard English names of amphibians and reptiles of North America north of Mexico, with comments regarding confidence in our understanding. 7th edition. SSAR Herpetological Circular 39:1-92.
  18. Crother, B. I. (editor). 2017. Scientific and standard English names of amphibians and reptiles of North America north of Mexico, with comments regarding confidence in our understanding. 8th edition. SSAR Herpetological Circular 43:1-104. [Updates in SSAR North American Species Names Database at: https://ssarherps.org/cndb]
  19. Dundee, H. A., and D. A. Rossman. 1989. The amphibians and reptiles of Louisiana. Louisiana State University Press, Baton Rouge.
  20. Ernst, C. H. 1992. Venomous reptiles of North America. Smithsonian Institution Press, Washington, D.C. ix + 236 pp.
  21. Ernst, C. H., and E. M. Ernst. 2003. Snakes of the United States and Canada. Smithsonian Books, Washington, D.C.
  22. Ernst, C. H., and R. W. Barbour. 1989b. Snakes of eastern North America. George Mason Univ. Press, Fairfax, Virginia. 282 pp.
  23. Fitch, H. S. 1960. Autecology of the copperhead. Univ. Kansas Pub. Mus. Nat. Hist. 13:85-288.
  24. Gibbons, J. W., and R. D. Semlitsch. 1991. Guide to the reptiles and amphibians of the Savannah River Site. Univ. of Georgia Press, Athens. xii + 131 pp.
  25. Global Biodiversity Information Facility (GBIF). 2024. Global Biodiversity Information Facility (GBIF) data portal. Online. Available: https://www.gbif.org/ (accessed 2024).
  26. Gloyd, H. K., and R. Conant. 1990. Snakes of the <i>Agkistrodon </i> complex: a monographic review. Society for the Study of Amphibians and Reptiles. vi + 614 pp.
  27. Green, N. B., and T. K. Pauley. 1987. Amphibians and reptiles in West Virginia. University of Pittsburgh Press, Pittsburgh, Pennsylvania. xi + 241 pp.
  28. Guiher, T.J. and F.T. Burbrink. 2008. Demographic and phylogeographic histories of two venomous North American snakes of the genus <i>Agkistrodon</i>. Molecular Phylogenetics and Evolution 48: 543–553.
  29. Hulse, A. C., C. J. McCoy, and E. Censky. 2001. Amphibians and reptiles of Pennsylvania and the Northeast. Comstock Publishing Associates, Cornell University Press, Ithaca. 419 pp.
  30. Johnson, T. R. 1987. The amphibians and reptiles of Missouri. Missouri Department of Conservation, Jefferson City. 368 pp.
  31. Johnson, T. R. 2000. The amphibians and reptiles of Missouri. Second edition. Missouri Department of Conservation, Jefferson City. 400 pp.
  32. Klemens, M. W. 1993. Amphibians and reptiles of Connecticut and adjacent regions. State Geological and Natural History Survey of Connecticut, Bulletin 112. xii + 318 pp.
  33. Martof, B. S., W. M. Palmer, J. R. Bailey, and J. R. Harrison, III. 1980. Amphibians and reptiles of the Carolinas and Virginia. University of North Carolina Press, Chapel Hill, North Carolina. 264 pp.
  34. Minton, S. A., Jr. 1972. Amphibians and reptiles of Indiana. Indiana Academy Science Monographs 3. v + 346 pp.
  35. Mitchell, J. C. 1994. The reptiles of Virginia. Smithsonian Institution Press, Washington, D.C. xv + 352 pp.
  36. Mount, R. H. 1975. The reptiles and amphibians of Alabama. Auburn University Agricultural Experiment Station, Auburn, Alabama. vii + 347 pp.
  37. NatureServe's Rapid Analysis of Rarity and Endangerment Conservation Assessment Tool (RARECAT). 2024. Version: 1.1.1 (released Oct 01, 2024).
  38. Palmer, W. M., and A. L. Braswell. 1995. Reptiles of North Carolina. North Carolina State Museum of Natural Sciences, University of North Carolina Press, Chapel Hill, North Carolina.
  39. Powell, R., R. Conant, and J. T. Collins. 2016. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America, fourth edition. Houghton Mifflin Harcourt, Boston and New York. 512 pp.
  40. Smith, P. W. 1961. The amphibians and reptiles of Illinois. Illinois Natural History Survey 28(1):1-298.
  41. Tennant, A. 1984. The Snakes of Texas. Texas Monthly Press, Austin, Texas. 561 pp.
  42. Trauth, S. E., H. W. Robison, and M. V. Plummer. 2004. The amphibians and reptiles of Arkansas. University of Arkansas Press.
  43. Webb, R. G. 1970. Reptiles of Oklahoma. University of Oklahoma Press, Norman. 370 pp.
  44. Werler, J. E., and J. R. Dixon. 2000. Texas snakes: identification, distribution, and natural history. University of Texas Press, Austin. xv + 437 pp.