Leucosticte atrata

Ridgeway, 1874

Black Rosy-Finch

G4Apparently Secure Found in 333 roadless areas NatureServe Explorer →
G4Apparently SecureGlobal Rank
EndangeredIUCN
Black Rosy-Finch (Leucosticte atrata). © Eric Konkol; Cornell Lab of Ornithology | Macaulay Library.
© Eric Konkol; Cornell Lab of Ornithology | Macaulay Library
Black Rosy-Finch (Leucosticte atrata). © Lesley Tullis; Cornell Lab of Ornithology | Macaulay Library.
© Lesley Tullis; Cornell Lab of Ornithology | Macaulay Library
Black Rosy-Finch (Leucosticte atrata). © Levi Plummer; Cornell Lab of Ornithology | Macaulay Library.
© Levi Plummer; Cornell Lab of Ornithology | Macaulay Library
Black Rosy-Finch (Leucosticte atrata). © Linda Chittum; Cornell Lab of Ornithology | Macaulay Library.
© Linda Chittum; Cornell Lab of Ornithology | Macaulay Library
Black Rosy-Finch (Leucosticte atrata). © Dorian Anderson; Cornell Lab of Ornithology | Macaulay Library.
© Dorian Anderson; Cornell Lab of Ornithology | Macaulay Library
Black Rosy-Finch (Leucosticte atrata). © Martina Nordstrand; Cornell Lab of Ornithology | Macaulay Library.
© Martina Nordstrand; Cornell Lab of Ornithology | Macaulay Library
Black Rosy-Finch (Leucosticte atrata). Photo by U.S. Fish & Wildlife Service, Public Domain (U.S. Government Work), via ECOS.
U.S. Fish & Wildlife Service, https://www.usa.gov/government-works
Identity
Unique IDELEMENT_GLOBAL.2.105480
Element CodeABPBY02010
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNEndangered
Endemicendemic to a single nation
KingdomAnimalia
PhylumCraniata
ClassAves
OrderPasseriformes
FamilyFringillidae
GenusLeucosticte
Other Common Names
black rosy-finch (EN) Roselin noir (FR)
Concept Reference
American Ornithologists' Union (AOU). 1998. Check-list of North American birds. Seventh edition. American Ornithologists' Union, Washington, D.C. [as modified by subsequent supplements and corrections published in The Auk]. Also available online: http://www.aou.org/.
Taxonomic Comments
Prior to 1983, North American rosy-finches were regarded as three species (L. atrata, L. australis, and L. tephrocotis). AOU (1983) lumped these together with Asian species as L. arctoa. Subsequently, Sibley and Monroe (1990, who cited unpublished genetic, biochemical, and morphological data by French and Loskot) and AOU (1993, who stated that the 1983 merger was based on insufficient new information) again recognized three species of rosy-finches in North America, distinct from Old World L. arctoa.

The three North American species sometimes have been merged as L. tephrocotis (American Rosy-Finch). Unpublished work by Johnson (1972) recognized a fourth North American species (L. griseonucha, comprising nominal subspecies griseonucha and umbrina of the Pribilof and Aleutian islands), but this taxon has not been accepted as a full species in subsequent checklists.
Conservation Status
Review Date1996-12-04
Change Date1996-12-04
Edition Date1995-03-08
Edition AuthorsR.E. Johnson and J.D. Reichel. Minor revisions by L. Master & G. Hammerson.
Range Extent20,000-2,500,000 square km (about 8000-1,000,000 square miles)
Number of Occurrences21 - 80
Rank Reasons
This species has moderate numbers of occurrences, limited range, and is somewhat numerous. However, approximately 30% of the population is concentrated in a single occurrence, the Beartooth Mountains. Most occurrences are on public lands, and development in alpine areas (e.g., mining) has apparently not caused population declines. Overall, it apparently secure.
Range Extent Comments
BREEDS: mountains from central Idaho, southwestern and south-central Montana, and northwestern and north-central Wyoming south to southeastern Oregon, northeastern and east-central Nevada (south to the Snake Mountains), and central Utah (to the Tushar and La Sal mountains). Beartooth Mounatins have more than 30% of the global population. WINTERS: central Idaho and western and southeastern Wyoming south to eastern California (at least casually), southern Nevada, northern Arizona, and northern New Mexico (AOU 1983).
Threat Impact Comments
Only known threat is new massive mining which could level many square miles of breeding habitat.
Ecology & Habitat

Habitat

Barren, rocky or grassy areas and cliffs among glaciers or beyond timberline; in migration and winter also in open situations, fields, cultivated lands, brushy areas, and around human habitation (AOU 1983). May roost in mine shaft or similar protected site. Nests usually in rock crevices or holes in cliffs above snow fields. May nest in old abandoned building.

Ecology

Males typically outnumber females in breeding and wintering populations. During breeding season male defends "territory" around female wherever she moves (Ryser 1985). Forms large flocks when not breeding.

Reproduction

Clutch size usually is 4-5. Incubation lasts 12-14 days, by female. Young are tended by both adults, leave nest at about 20 days.
Terrestrial Habitats
Shrubland/chaparralGrassland/herbaceousAlpineBare rock/talus/screeCliffCropland/hedgerow
Other Nations (1)
United StatesN4
ProvinceRankNative
Navajo NationS1NYes
OregonS2BYes
MontanaS2Yes
WyomingS1B,S2NYes
UtahS2Yes
NevadaS2Yes
ColoradoS4NYes
New MexicoS3NYes
IdahoS2Yes
Roadless Areas (333)
California (6)
AreaForestAcres
ButtermilkInyo National Forest542
ButtermilkInyo National Forest542
North LakeInyo National Forest2,406
North LakeInyo National Forest2,406
Table Mtn.Inyo National Forest4,215
Table Mtn.Inyo National Forest4,215
Colorado (9)
AreaForestAcres
Bristol HeadRio Grande NF46,087
Bristol HeadRio Grande NF46,087
HermosaSan Juan NF148,103
HermosaSan Juan NF148,103
HermosaSan Juan NF148,103
San MiguelSan Juan NF64,263
San MiguelSan Juan NF64,263
Williams Fork Ptarmingan AdjacentArapaho & Roosevelt NFs36,351
Williams Fork Ptarmingan AdjacentArapaho & Roosevelt NFs36,351
Idaho (20)
AreaForestAcres
Borah PeakSalmon-Challis National Forest130,463
Borah PeakSalmon-Challis National Forest130,463
Borah PeakSalmon-Challis National Forest130,463
Boulder - White CloudsSawtooth National Forest322,653
Boulder - White CloudsSalmon-Challis National Forest139,297
Boulder - White CloudsSalmon-Challis National Forest139,297
Boulder - White CloudsSawtooth National Forest322,653
Italian PeakCaribou-Targhee National Forest141,158
Italian PeakCaribou-Targhee National Forest141,158
Lemhi RangeSalmon-Challis National Forest308,533
Lemhi RangeSalmon-Challis National Forest308,533
PalisadesCaribou-Targhee National Forest122,002
PalisadesCaribou-Targhee National Forest122,002
PalisadesCaribou-Targhee National Forest122,002
Pioneer MountainsSalmon-Challis National Forest172,460
Pioneer MountainsSalmon-Challis National Forest172,460
Pioneer MountainsSawtooth National Forest119,563
Pioneer MountainsSawtooth National Forest119,563
Pioneer MountainsSawtooth National Forest119,563
Pioneer MountainsSalmon-Challis National Forest172,460
Montana (52)
AreaForestAcres
BeartoothGallatin National Forest5,285
BeartoothGallatin National Forest5,285
Black ButteCuster National Forest871
Black ButteCuster National Forest871
BridgerGallatin National Forest45,059
BridgerGallatin National Forest45,059
Cabin Creek Wildlife Management Area OcdGallatin National Forest35,048
Cabin Creek Wildlife Management Area OcdGallatin National Forest35,048
Chico PeakGallatin National Forest10,744
Chico PeakGallatin National Forest10,744
Crazy MountainGallatin National Forest82,093
Crazy MountainGallatin National Forest82,093
East PioneerBeaverhead-Deerlodge National Forest145,082
East PioneerBeaverhead-Deerlodge National Forest145,082
Gallatin FringeGallatin National Forest51,571
Gallatin FringeGallatin National Forest51,571
Garfield MountainBeaverhead-Deerlodge National Forest41,891
Garfield MountainBeaverhead-Deerlodge National Forest41,891
Hyalite - Porcupine - Buffalo Horn Wilderness Study AreaGallatin National Forest143,991
Hyalite - Porcupine - Buffalo Horn Wilderness Study AreaGallatin National Forest143,991
Italian PeakBeaverhead-Deerlodge National Forest90,401
Italian PeakBeaverhead-Deerlodge National Forest90,401
Line Creek PlateauCuster National Forest24,825
Line Creek PlateauCuster National Forest24,825
Line Creek PlateauCuster National Forest24,825
LionheadGallatin National Forest33,549
LionheadGallatin National Forest33,549
Middle Mtn. / Tobacco RootsBeaverhead-Deerlodge National Forest96,487
Middle Mtn. / Tobacco RootsBeaverhead-Deerlodge National Forest96,487
Mt. Gmt Area HCuster National Forest1,335
Mt. Gmt Area HCuster National Forest1,335
Mt. Gmt Area HCuster National Forest1,335
North AbsarokaGallatin National Forest159,075
North AbsarokaCuster National Forest21,063
North AbsarokaCuster National Forest21,063
North AbsarokaCuster National Forest21,063
North AbsarokaGallatin National Forest159,075
Proposed Line Creek PrnaCuster National Forest389
Proposed Line Creek PrnaCuster National Forest389
Proposed Line Creek PrnaCuster National Forest389
Red Lodge Creek HellroaringCuster National Forest17,210
Red Lodge Creek HellroaringCuster National Forest17,210
ReefGallatin National Forest2,500
ReefGallatin National Forest2,500
Republic MountainGallatin National Forest836
Republic MountainGallatin National Forest836
Rock CreekCuster National Forest100
Rock CreekCuster National Forest100
Selway - Bitterroot (01067)Bitterroot National Forest114,953
Selway - Bitterroot (01067)Bitterroot National Forest114,953
West Big HoleBeaverhead-Deerlodge National Forest133,563
West Big HoleBeaverhead-Deerlodge National Forest133,563
Nevada (27)
AreaForestAcres
Bald Mtn.Humboldt-Toiyabe National Forest41,598
Bald Mtn.Humboldt-Toiyabe National Forest41,598
Humboldt - Angel LkHumboldt-Toiyabe National Forest1,008
Humboldt - Angel LkHumboldt-Toiyabe National Forest1,008
Humboldt - LizziesHumboldt-Toiyabe National Forest70
Humboldt - LizziesHumboldt-Toiyabe National Forest70
Humboldt - Trout CrHumboldt-Toiyabe National Forest1,346
Humboldt - Trout CrHumboldt-Toiyabe National Forest1,346
Humboldt - WinchellHumboldt-Toiyabe National Forest1,272
Humboldt - WinchellHumboldt-Toiyabe National Forest1,272
Ruby - Horse Mtn.Humboldt-Toiyabe National Forest613
Ruby - Horse Mtn.Humboldt-Toiyabe National Forest613
Ruby - Lamoille CynHumboldt-Toiyabe National Forest32,771
Ruby - Lamoille CynHumboldt-Toiyabe National Forest32,771
Ruby - Lamoille CynHumboldt-Toiyabe National Forest32,771
Ruby - MarshHumboldt-Toiyabe National Forest10,164
Ruby - MarshHumboldt-Toiyabe National Forest10,164
Ruby - ThompsonHumboldt-Toiyabe National Forest5,289
Ruby - ThompsonHumboldt-Toiyabe National Forest5,289
Snake - Mt. WashingtonHumboldt-Toiyabe National Forest3,749
Snake - Mt. WashingtonHumboldt-Toiyabe National Forest3,749
Snake - Peacock CynHumboldt-Toiyabe National Forest1,069
Snake - Peacock CynHumboldt-Toiyabe National Forest1,069
Snake - Pole CkHumboldt-Toiyabe National Forest79
Snake - Pole CkHumboldt-Toiyabe National Forest79
Snake - Pole CynHumboldt-Toiyabe National Forest766
Snake - Pole CynHumboldt-Toiyabe National Forest766
New Mexico (27)
AreaForestAcres
Bull - Of - The - WoodsCarson National Forest488
Bull - Of - The - WoodsCarson National Forest488
Canones CreekSanta Fe National Forest3,938
Canones CreekSanta Fe National Forest3,938
Columbine - Hondo Wilderness Study AreaCarson National Forest43,739
Columbine - Hondo Wilderness Study AreaCarson National Forest43,739
Columbine - Hondo Wilderness Study AreaCarson National Forest43,739
Cruces BasinCarson National Forest5,244
Cruces BasinCarson National Forest5,244
Ghost TownSanta Fe National Forest219
Ghost TownSanta Fe National Forest219
Guaje CanyonSanta Fe National Forest6,104
Guaje CanyonSanta Fe National Forest6,104
Latir PeakCarson National Forest3,573
Latir PeakCarson National Forest3,573
Oser MesaCarson National Forest2,840
Oser MesaCarson National Forest2,840
Peralta RidgeSanta Fe National Forest4,027
Peralta RidgeSanta Fe National Forest4,027
PolvaderaSanta Fe National Forest2,487
PolvaderaSanta Fe National Forest2,487
Polvadera PeakSanta Fe National Forest6,291
Polvadera PeakSanta Fe National Forest6,291
RendijaSanta Fe National Forest2,176
RendijaSanta Fe National Forest2,176
Wheeler Peak WildernessCarson National Forest2,677
Wheeler Peak WildernessCarson National Forest2,677
Utah (88)
AreaForestAcres
0401001Ashley National Forest11,705
0401001Ashley National Forest11,705
0401009Ashley National Forest30,378
0401009Ashley National Forest30,378
0401010Ashley National Forest21,886
0401010Ashley National Forest21,886
0401016Ashley National Forest5,695
0401016Ashley National Forest5,695
0401023Ashley National Forest8,352
0401023Ashley National Forest8,352
0401024Ashley National Forest12,882
0401024Ashley National Forest12,882
0401027Ashley National Forest7,312
0401027Ashley National Forest7,312
0401031Ashley National Forest7,110
0401031Ashley National Forest7,110
0401032Ashley National Forest6,471
0401032Ashley National Forest6,471
0419020Ashley National Forest355,684
0419020Ashley National Forest355,684
0419020Ashley National Forest355,684
418001Uinta National Forest5,697
418001Uinta National Forest5,697
418016Uinta National Forest35,240
418016Uinta National Forest35,240
418017Uinta National Forest19,631
418017Uinta National Forest19,631
418019Uinta National Forest6,854
418019Uinta National Forest6,854
418024Uinta National Forest51,699
418024Uinta National Forest51,699
418029Uinta National Forest15,673
418029Uinta National Forest15,673
418040Uinta National Forest1,702
418040Uinta National Forest1,702
Boulder Mtn. / Boulder Top / Deer LakeDixie National Forest110,690
Boulder Mtn. / Boulder Top / Deer LakeDixie National Forest110,690
Dairy ForkManti-Lasal National Forest30,222
Dairy ForkManti-Lasal National Forest30,222
FishhookDixie National Forest12,959
FishhookDixie National Forest12,959
FishhookDixie National Forest12,959
GibsonWasatch-Cache National Forest5,350
GibsonWasatch-Cache National Forest5,350
High Uintas (UT)Wasatch-Cache National Forest102,398
High Uintas (UT)Wasatch-Cache National Forest102,398
High Uintas (UT)Wasatch-Cache National Forest102,398
HogsbackWasatch-Cache National Forest7,936
HogsbackWasatch-Cache National Forest7,936
Horse Mountain - Mans PeakManti-Lasal National Forest22,159
Horse Mountain - Mans PeakManti-Lasal National Forest22,159
LakesWasatch-Cache National Forest121,967
LakesWasatch-Cache National Forest121,967
LakesWasatch-Cache National Forest121,967
Lewis PeakWasatch-Cache National Forest11,616
Lewis PeakWasatch-Cache National Forest11,616
Lone Peak ContiguousWasatch-Cache National Forest874
Lone Peak ContiguousWasatch-Cache National Forest874
Lone Peak ContiguousWasatch-Cache National Forest874
Long Neck Mesa / Steep Creek / Oak Creek - Steep Creek / OakDixie National Forest55,489
Long Neck Mesa / Steep Creek / Oak Creek - Steep Creek / OakDixie National Forest55,489
Mt. Logan NorthWasatch-Cache National Forest18,930
Mt. Logan NorthWasatch-Cache National Forest18,930
Mt. NaomiWasatch-Cache National Forest41,922
Mt. NaomiWasatch-Cache National Forest41,922
Mt. OlympusWasatch-Cache National Forest9,982
Mt. OlympusWasatch-Cache National Forest9,982
Mt. OlympusWasatch-Cache National Forest9,982
New Home BenchDixie National Forest10,513
New Home BenchDixie National Forest10,513
NoblettsWasatch-Cache National Forest3,116
NoblettsWasatch-Cache National Forest3,116
Public GroveWasatch-Cache National Forest6,341
Public GroveWasatch-Cache National Forest6,341
South FrancisWasatch-Cache National Forest3,374
South FrancisWasatch-Cache National Forest3,374
Swan Creek MountainWasatch-Cache National Forest9,390
Swan Creek MountainWasatch-Cache National Forest9,390
Temple PeakWasatch-Cache National Forest24,081
Temple PeakWasatch-Cache National Forest24,081
Twin PeaksWasatch-Cache National Forest6,157
Twin PeaksWasatch-Cache National Forest6,157
Twin PeaksWasatch-Cache National Forest6,157
WellsvilleWasatch-Cache National Forest1,717
WellsvilleWasatch-Cache National Forest1,717
White PineWasatch-Cache National Forest1,942
White PineWasatch-Cache National Forest1,942
White PineWasatch-Cache National Forest1,942
Wyoming (104)
AreaForestAcres
Beartooth Proposed WildernessShoshone National Forest16,837
Beartooth Proposed WildernessShoshone National Forest16,837
Beartooth Proposed WildernessShoshone National Forest16,837
Campbell LakeMedicine Bow-Routt National Forest7,088
Campbell LakeMedicine Bow-Routt National Forest7,088
Canyon CreekShoshone National Forest7,530
Canyon CreekShoshone National Forest7,530
Cloud Peak ContiguousBighorn National Forest113,757
Cloud Peak ContiguousBighorn National Forest113,757
Cloud Peak ContiguousBighorn National Forest113,757
Devils CanyonBighorn National Forest37,416
Devils CanyonBighorn National Forest37,416
Dunoir Special Management UnitShoshone National Forest29,719
Dunoir Special Management UnitShoshone National Forest29,719
East DunoirShoshone National Forest3,175
East DunoirShoshone National Forest3,175
Franc's PeakShoshone National Forest68,561
Franc's PeakShoshone National Forest68,561
French CreekMedicine Bow-Routt National Forest5,928
French CreekMedicine Bow-Routt National Forest5,928
Grayback RidgeBridger-Teton National Forest295,113
Grayback RidgeBridger-Teton National Forest295,113
Gros Ventre MountainsBridger-Teton National Forest106,418
Gros Ventre MountainsBridger-Teton National Forest106,418
Gros Ventre MountainsBridger-Teton National Forest106,418
Hideout CreekBighorn National Forest10,096
Hideout CreekBighorn National Forest10,096
Horse Creek MesaBighorn National Forest77,808
Horse Creek MesaBighorn National Forest77,808
Libby FlatsMedicine Bow-Routt National Forest11,107
Libby FlatsMedicine Bow-Routt National Forest11,107
Little BighornBighorn National Forest133,949
Little BighornBighorn National Forest133,949
Little GooseBighorn National Forest25,558
Little GooseBighorn National Forest25,558
Little Popo AgieShoshone National Forest8,333
Little Popo AgieShoshone National Forest8,333
Little Sheep MountainBridger-Teton National Forest14,192
Little Sheep MountainBridger-Teton National Forest14,192
Little Sheep MountainBridger-Teton National Forest14,192
Middle ForkShoshone National Forest51,772
Middle ForkShoshone National Forest51,772
Middle ForkShoshone National Forest51,772
Middle ForkMedicine Bow-Routt National Forest13,238
Middle ForkMedicine Bow-Routt National Forest13,238
Middle ForkMedicine Bow-Routt National Forest13,238
Mosquito Lake - Seven LakesBridger-Teton National Forest51,950
Mosquito Lake - Seven LakesBridger-Teton National Forest51,950
Mosquito Lake - Seven LakesBridger-Teton National Forest51,950
Munger MountainBridger-Teton National Forest12,827
Munger MountainBridger-Teton National Forest12,827
North BoundaryShoshone National Forest182
North BoundaryShoshone National Forest182
Pacific Creek - Blackrock CreekBridger-Teton National Forest24,658
Pacific Creek - Blackrock CreekBridger-Teton National Forest24,658
PalisadesTarghee National Forest1,121
PalisadesTarghee National Forest1,121
PalisadesTarghee National Forest1,121
Pass CreekShoshone National Forest6,172
Pass CreekShoshone National Forest6,172
Phillips RidgeBridger-Teton National Forest10,108
Phillips RidgeBridger-Teton National Forest10,108
Phillips RidgeBridger-Teton National Forest10,108
ReefShoshone National Forest16,817
ReefShoshone National Forest16,817
Snowy RangeMedicine Bow-Routt National Forest29,660
Snowy RangeMedicine Bow-Routt National Forest29,660
South Beartooth HighwayShoshone National Forest105,570
South Beartooth HighwayShoshone National Forest105,570
South Beartooth HighwayShoshone National Forest105,570
South DunoirShoshone National Forest2,749
South DunoirShoshone National Forest2,749
South ForkShoshone National Forest64,903
South ForkShoshone National Forest64,903
South Wyoming RangeBridger-Teton National Forest85,776
South Wyoming RangeBridger-Teton National Forest85,776
Spread Creek - Gros Ventre RiverBridger-Teton National Forest166,097
Spread Creek - Gros Ventre RiverBridger-Teton National Forest166,097
Sulphur CreekShoshone National Forest30,221
Togwotee PassShoshone National Forest7,344
Togwotee PassShoshone National Forest7,344
Trout CreekShoshone National Forest44,034
Trout CreekShoshone National Forest44,034
Walker PrairieBighorn National Forest62,434
Walker PrairieBighorn National Forest62,434
Wapiti Valley NorthShoshone National Forest18,345
Wapiti Valley NorthShoshone National Forest18,345
Wapiti Valley SouthShoshone National Forest43,517
Wapiti Valley SouthShoshone National Forest43,517
West DunoirShoshone National Forest2,542
West DunoirShoshone National Forest2,542
West Slope TetonsTarghee National Forest47,448
West Slope TetonsTarghee National Forest47,448
West Slope TetonsTarghee National Forest47,448
West Slope WindsBridger-Teton National Forest143,252
West Slope WindsBridger-Teton National Forest143,252
West Slope WindsBridger-Teton National Forest143,252
Wilderness Study AreaTarghee National Forest51,961
Wilderness Study AreaTarghee National Forest51,961
Windy MountainShoshone National Forest31,283
Windy MountainShoshone National Forest31,283
Windy MountainShoshone National Forest31,283
Wood RiverShoshone National Forest50,638
Wood RiverShoshone National Forest50,638
References (14)
  1. American Ornithologists' Union (AOU). 1983. Check-list of North American Birds, 6th edition. Allen Press, Inc., Lawrence, Kansas. 877 pp.
  2. American Ornithologists' Union (AOU). 1993. Thirty-ninth supplement to the American Ornithologists' Union Check-list of North American Birds. Auk 110:675-82.
  3. American Ornithologists' Union (AOU). 1998. Check-list of North American birds. Seventh edition. American Ornithologists' Union, Washington, D.C. [as modified by subsequent supplements and corrections published in <i>The Auk</i>]. Also available online: http://www.aou.org/.
  4. Balda, R. P., and G. C. Bateman. 1971. Flocking and annual cycle of the piñon jay, <i>Gymnorhinus cyanocephalus</i>. Condor 73:287-302.
  5. Horn, H. S. 1968. The adaptive significance of colonial nesting in the Brewer's Blackbird. Ecology 49:682-694.
  6. Johnson, R. E. 1972e. The biosystematics of the avian genus <i>Leucosticte</i>. Ph.D dissertation. University of California, Berkeley, California, USA.
  7. Ligon, J. D. 1971. Late summer-autumnal breeding of the piñon jay in New Mexico. Condor 73:147-153.
  8. Moore, W. S., and R. A. Dolbeer. 1989. The use of banding recovery data to estimate dispersal rates and gene flow in avian species: case studies in the Red-winged Blackbird and Common Grackle. Condor 91:242-253.
  9. Sibley, C.G., and B.L. Monroe, Jr. 1990. Distribution and Taxonomy of Birds of the World. Yale University Press, New Haven, CT. xxiv + 1111 pp.
  10. Sibley, C. G., and J. E. Ahlquist. 1990. Phylogeny and classification of birds: a study in molecular evolution. Yale Univ. Press, New Haven. xxiii + 976 pp.
  11. Tarvin, K. A., and G. E. Woolfenden. 1999. Blue Jay (<i>Cyanocitta cristata</i>). No. 469 IN A. Poole and F. Gill, editors. The birds of North America. The Birds of North America, Inc., Philadelphia, PA. 32pp.
  12. Thompson, F. R., III. 1994. Temporal and spatial patterns of breeding brown-headed cowbirds in the midwestern United States. Auk 111:979-990.
  13. Williams, L. 1952b. Breeding behavior of the Brewer blackbird. Condor 54:3-47.
  14. Willson, M. F. 1966. Breeding ecology of the Yellow-headed Blackbird. Ecological Monographs 36:51-77.