Tsuga canadensis

(L.) Carr.

Eastern Hemlock

G4Apparently Secure (G4G5) Found in 135 roadless areas NatureServe Explorer →
G4Apparently SecureGlobal Rank
Near threatenedIUCN
High - mediumThreat Impact
Eastern Hemlock (Tsuga canadensis). Photo by Joel Stave, CC BY-NC 4.0, via iNaturalist.
Joel Stave, CC BY-NC 4.0
Eastern Hemlock (Tsuga canadensis). Photo by aaronbuffington, CC BY-NC 4.0, via iNaturalist.
aaronbuffington, CC BY-NC 4.0
Eastern Hemlock (Tsuga canadensis). Photo by agbelliveau, CC0 1.0, via iNaturalist.
agbelliveau, CC0 1.0
Eastern Hemlock (Tsuga canadensis). Photo by agbelliveau, CC0 1.0, via iNaturalist.
agbelliveau, CC0 1.0
Identity
Unique IDELEMENT_GLOBAL.2.131718
Element CodePGPIN06010
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVascular Plant
IUCNNear threatened
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomPlantae
PhylumConiferophyta
ClassPinopsida
OrderPinales
FamilyPinaceae
GenusTsuga
Other Common Names
eastern hemlock (EN) Pruche du Canada (FR)
Concept Reference
Kartesz, J.T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. 2nd edition. 2 vols. Timber Press, Portland, OR.
Conservation Status
Rank Method Rank calculation - Biotics v2
Review Date2021-06-29
Change Date2021-06-29
Edition Date2020-04-29
Edition AuthorsThurman, Carolyn (1993), rev. L. Morse (1995), rev. Adele Olivero (2002), rev. Treher (2020)
Threat ImpactHigh - medium
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences> 300
Rank Reasons
Tsuga canadensis occurs from Ontario to Nova Scotia south to Maryland and eastern Minnesota, and along the Appalachian mountains to Georgia and Alabama. This species has declined, most severely in the southern Appalachians, due to mortality caused by the Hemlock Wooly Adelgid (Adelges tsugae), an exotic pest. The Conservation Status of this species should be reviewed frequently to detect changes in the impact of the pest.
Range Extent Comments
Tsuga canadensis occurs from Ontario to Nova Scotia south to Maryland and eastern Minnesota, and along the Appalachian mountains to Georgia and Alabama (Fernald 1950, Kartesz 1999).
Threat Impact Comments
A serious threat to the species is the hemlock woolly adelgid (Adelges tsugae (Annand, 1928)), an insect that is believed to have originated in Japan and first detected in North America in the 1950s. The adelgid, found in 19 states and 2 provinces, has caused mortality in Canadian and Carolina hemlocks throughout the Appalachian region, especially in the south, but also as far north as southern Maine, Michigan, Nova Scotia, and Ontario (Ellison et al. 2018, Cohn 1993). The most serious declines are found in the southern Appalachian Mountains, areas of the mid-Atlantic region, and southern New England (Ellison et al. 2018). The adelgid is a chronic problem, as it can infect all ages of trees. Trees that are attacked can die in several years, but most die slowly over a decade. Mortality is variable, from 0 to 99% (Ellison et al. 2018, Souto and Shields 1999). A related threat is "preemptive logging" which may be occurring well outside the insect's present range (Foster 1999). In Canada, the spread of the adelgid is being carefully monitored; when it is found, eradication is attempted (Fidgen et al. 2015). Nova Scotia is experiencing declines due to the adelgid (Emilson and Stastny 2019). The habitat shift after infestation to deciduous hardwoods, as well as pre-emptive logging to prevent spread of the adelgid, which also causes a habitat shift, contribute to an environment that is not conducive to the regeneration of the species.

In addition to the adelgid, the non-native Elongate Hemlock Scale is also stressing the species (Miller-Pierce et al. 2019). The introduced Spongy Moth (Lymantria dispar) also attack hemlocks, and a recent study has shown that Spongy Moths feed preferentially on branches and leaves infected with adelgid (Kinahan et al. 2019).
Ecology & Habitat

Habitat

Mostly hilly or rocky woods (Fernald, 1950); in southern portion of range, frequently in moist, shaded ravines, or other sheltered microhabitats.
Other Nations (2)
United StatesN4
ProvinceRankNative
KentuckyS5Yes
New JerseyS4Yes
TennesseeSNRYes
MaineS5Yes
ConnecticutSNRYes
GeorgiaSNRYes
MichiganSNRYes
North CarolinaS4Yes
AlabamaS4Yes
WisconsinSNRYes
South CarolinaS3Yes
West VirginiaS3Yes
MarylandSNRYes
MinnesotaS1Yes
DelawareS1Yes
OhioSNRYes
PennsylvaniaS5Yes
IndianaS3Yes
VirginiaS4Yes
VermontS5Yes
MassachusettsSNRYes
New YorkS5Yes
Rhode IslandSNRYes
New HampshireSNRYes
CanadaN5
ProvinceRankNative
QuebecS4Yes
Nova ScotiaS4Yes
Prince Edward IslandS3Yes
New BrunswickS5Yes
OntarioS5Yes
Threat Assessments
ThreatScopeSeverityTiming
8 - Invasive & other problematic species, genes & diseasesLarge - restrictedSerious or 31-70% pop. declineHigh (continuing)
8.1 - Invasive non-native/alien species/diseasesLarge - restrictedSerious or 31-70% pop. declineHigh (continuing)
8.1.2 - Named speciesLarge - restrictedSerious or 31-70% pop. declineHigh (continuing)

Plant Characteristics
Economic Value (Genus)No
Roadless Areas (135)
Georgia (13)
AreaForestAcres
Boggs CreekChattahoochee National Forest2,073
Cedar MountainChattahoochee National Forest1,083
Ellicott Rock AdditionChattahoochee National Forest690
Helton CreekChattahoochee National Forest2,348
Indian Grave GapChattahoochee National Forest1,020
Joe GapChattahoochee National Forest5,321
Kelly RidgeChattahoochee National Forest8,325
Ken MountainChattahoochee National Forest527
Lance CreekChattahoochee National Forest9,025
Miller CreekChattahoochee National Forest701
Pink KnobChattahoochee National Forest12,127
Rocky MountainChattahoochee National Forest4,269
Sarah's CreekChattahoochee National Forest6,888
Kentucky (1)
AreaForestAcres
WolfpenDaniel Boone National Forest2,835
Maine (1)
AreaForestAcres
Caribou - Speckled ExtWhite Mountain National Forest5,988
New Hampshire (12)
AreaForestAcres
Carr MountainWhite Mountain National Forest17,110
Dartmouth RangeWhite Mountain National Forest9,233
Great Gulf Ext.White Mountain National Forest15,110
JobildunkWhite Mountain National Forest3,660
Kinsman MountainWhite Mountain National Forest8,999
Mt. Wolf - Gordon PondWhite Mountain National Forest11,846
PemigewassetWhite Mountain National Forest32,255
Pemigewasset ExtWhite Mountain National Forest15,840
Presidential - Dry River ExtWhite Mountain National Forest10,555
Sandwich RangeWhite Mountain National Forest16,797
WatervilleWhite Mountain National Forest4,312
Wild RiverWhite Mountain National Forest46,878
North Carolina (26)
AreaForestAcres
Bald MountainPisgah National Forest11,085
Balsam ConePisgah National Forest10,591
Barkers Creek (addition)Nantahala National Forest975
BearwallowPisgah National Forest4,113
Big Indian (addition)Nantahala National Forest1,155
Cheoah BaldNantahala National Forest7,795
Chunky Gal (addition)Nantahala National Forest3,336
Craggy MountainPisgah National Forest2,657
Dobson KnobPisgah National Forest6,111
Graveyard Ridge (addition)Pisgah National Forest1,958
Harper CreekPisgah National Forest7,325
Jarrett CreekPisgah National Forest7,485
Laurel MountainPisgah National Forest5,683
Linville Gorge AdditionPisgah National Forest2,809
Lost CovePisgah National Forest5,944
Mackey MountainPisgah National Forest5,934
Middle Prong AdditionPisgah National Forest1,852
Overflow CreekNantahala National Forest3,379
Sam Knob (addition)Pisgah National Forest2,576
Slide HollowPisgah National Forest193
SnowbirdNantahala National Forest8,489
South Mills RiverPisgah National Forest8,588
Tusquitee BaldNantahala National Forest13,670
Wilson CreekPisgah National Forest4,863
Woods MountainPisgah National Forest9,602
Yellowhammer Branch (add.)Nantahala National Forest1,255
Pennsylvania (5)
AreaForestAcres
Allegheny FrontAllegheny National Forest7,430
CornplanterAllegheny National Forest2,929
Hearts ContentAllegheny National Forest221
Minister ValleyAllegheny National Forest1,417
Tracy RidgeAllegheny National Forest9,034
South Carolina (2)
AreaForestAcres
Ellicott Rock 1Sumter National Forest301
Ellicott Rock 2Sumter National Forest517
Tennessee (13)
AreaForestAcres
Bald MountainCherokee National Forest11,743
Bald River Gorge AdditionCherokee National Forest1,728
Beaver Dam CreekCherokee National Forest5,070
Brushy RidgeCherokee National Forest7,469
Devil's BackboneCherokee National Forest4,287
Flint Mill GapCherokee National Forest9,494
London Bridge BranchCherokee National Forest3,387
Rogers RidgeCherokee National Forest4,738
Sampson Mountain AdditionCherokee National Forest3,064
Slide HollowCherokee National Forest4,057
Stone MountainCherokee National Forest5,367
Sycamore CreekCherokee National Forest6,984
Upper Bald RiverCherokee National Forest9,202
Vermont (5)
AreaForestAcres
Bread LoafGreen Mountain and Finger Lakes National Forests1,768
Devil's Den 09083Green Mountain and Finger Lakes National Forests9,169
Griffith Lake 09084Green Mountain and Finger Lakes National Forests1,833
Wilder Mountain 09082Green Mountain and Finger Lakes National Forests8,759
Woodford 09086Green Mountain and Finger Lakes National Forests2,456
Virginia (37)
AreaForestAcres
Adams PeakGeorge Washington National Forest7,135
Bear CreekJefferson National Forest18,274
Beards MountainGeorge Washington National Forest7,505
Beartown Addition BJefferson National Forest2,985
Beaver Dam CreekJefferson National Forest1,135
Broad RunJefferson National Forest10,971
Brush MountainJefferson National Forest6,002
Brush Mountain EastJefferson National Forest4,916
Brushy MountainJefferson National Forest4,168
Dolly AnnGeorge Washington National Forest7,855
Elliott KnobGeorge Washington National Forest9,380
Gum RunGeorge Washington National Forest12,620
Hoop HoleJefferson National Forest4,652
Hunting Camp Little Wolf CreekJefferson National Forest8,953
JerkemtightGeorge Washington National Forest16,687
Kelley MountainGeorge Washington National Forest7,590
Laurel ForkGeorge Washington National Forest9,967
Lewis Fork AdditionJefferson National Forest749
Little AlleghanyGeorge Washington National Forest10,215
Little RiverGeorge Washington National Forest27,292
Mill MountainGeorge Washington National Forest10,840
Mountain Lake Addition AJefferson National Forest1,469
Mt. PleasantGeorge Washington National Forest8,933
New London Bridge BranchJefferson National Forest844
North MountainJefferson National Forest8,377
Northern MassanuttenGeorge Washington National Forest9,444
Oak KnobGeorge Washington National Forest10,882
Peters Mountain Addition BJefferson National Forest2,909
Raccoon BranchJefferson National Forest4,388
Ramseys Draft AdditionGeorge Washington National Forest12,781
Saint Marys AdditionGeorge Washington National Forest1,454
Seng MountainJefferson National Forest6,428
SkidmoreGeorge Washington National Forest5,641
Southern MassanuttenGeorge Washington National Forest11,985
The PriestGeorge Washington National Forest5,737
Three RidgesGeorge Washington National Forest4,745
Three SistersGeorge Washington National Forest8,149
West Virginia (17)
AreaForestAcres
Canaan LoopMonongahela National Forest7,867
Cheat MountainMonongahela National Forest8,191
Cranberry AdditionMonongahela National Forest11,123
Cranberry Glades Botanical AreaMonongahela National Forest785
Dolly Sods Roaring PlainMonongahela National Forest13,392
Dry River (WV)George Washington National Forest7,331
East Fork Of GreenbrierMonongahela National Forest7,167
Falls Of Hills CreekMonongahela National Forest6,925
Glady ForkMonongahela National Forest3,239
Little MountainMonongahela National Forest8,172
Marlin MountainMonongahela National Forest9,344
Mcgowan MountainMonongahela National Forest10,504
Middle MountainMonongahela National Forest19,020
North Mountain HopevilleMonongahela National Forest6,525
Seneca CreekMonongahela National Forest22,287
Tea Creek MountainMonongahela National Forest8,295
Turkey MountainMonongahela National Forest6,421
Wisconsin (3)
AreaForestAcres
09154 - St. Peters DomeChequamegon-Nicolet National Forest4,002
09159 - ThornappleChequamegon-Nicolet National Forest9,744
09164 - Tea LakeChequamegon-Nicolet National Forest5,510
References (16)
  1. Cohn, D'Vera. 1993. Shenandoah hemlocks under seige. Washington Post: September 4, 1993.
  2. Dharmadi, S.N., K.J. Elliott, C.F.Miniat. 2019. Lack of forest tree seedling recruitment and enhanced tree and shrub growth characterizes post-<i>Tsuga canadensis</i> mortality forests in the southern Appalachians. Forest Ecology and Management 440:122-130.
  3. Ellison, A.M., D.A. Orwig, M.C. Fitzpatrick, and E.L. Preisser. 2018. The Past, Present, and Future of the Hemlock Woolly Adelgid (<i>Adelges tsugae</i>) and Its Ecological Interactions with Eastern Hemlock (<i>Tsuga canadensis</i>) Forest. Insects 9:172.
  4. Emilson, C.E., and M. Stastny. 2019. A decision framework for hemlock woolly adelgid management: Review of the most suitable strategies and tactics for eastern Canada. Forest Ecology and Management 444:327-343.
  5. Fernald, M. L. 1950. Gray's manual of botany. 8th edition. Corrected printing (1970). D. Van Nostrand Company, New York. 1632 pp.
  6. Fidgen, J.G., M.C.Whitmore, and J.J. Turgeon. 2015. Detection of Hemlock Woolly Adelgid (Hemiptera: Adelgidae) Infestations with Sticky Traps. The Great Lakes Entomologist 48(3):125-131.
  7. Flora of North America Editorial Committee (FNA). 1993a. Flora of North America north of Mexico. Vol. 2. Pteridophytes and gymnosperms. Oxford Univ. Press, New York. xvi + 475 pp.
  8. Foster, D.R. 1999. Hemlock's future in the context of its history: An ecological perspective. In: McManus, K.A., K.S. Shields, and D.R. Souto eds. Proceedings: Symposium on Sustainable Management of Hemlock Ecosystems in Eastern North America, June 22-24, 1999. U.S. Forest Service, Northeastern Forest Experiment Station, General Technical Report NE-267. Durham, NH: 1-4.
  9. Kartesz, J.T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. 2nd edition. 2 vols. Timber Press, Portland, OR.
  10. Kartesz, J.T. 1999. A synonymized checklist and atlas with biological attributes for the vascular flora of the United States, Canada, and Greenland. First edition. In: Kartesz, J.T., and C.A. Meacham. Synthesis of the North American Flora, Version 1.0. North Carolina Botanical Garden, Chapel Hill, N.C.
  11. Kinahan, I.G., A.K. Baranowski, E.R. Whitney, S.K. Savage, C.M. Rigsby, E.E. Shoemaker, C.M. Orians, and E.L. Preisser. 2019 (online). Facilitation between invasive herbivores: hemlock woolly adelgid increases gypsy moth preference for and performance on eastern hemlock. Ecological Entomology https://doi.org/10.1111/een.12829.
  12. Little, E.L., Jr. 1979. Checklist of United States trees (native and naturalized). Agriculture Handbook No. 541. U.S. Forest Service, Washington, D.C. 375 pp.
  13. Miller-Pierce, M.R., D.A. Orwig, E. Preisser. 2019. Effects of Hemlock Woolly Adelgid and Elongate Hemlock Scale on Eastern Hemlock Growth and Foliar Chemistry, Environmental Entomology 39(2):513–519.
  14. Onken, B. P. 2001. Hemlock woolly adelgid - A race in time. U.S.D.A. Forest Service, Forest Health Protection, Morgantown, WV.
  15. Souto, D.R. and K.S. Shields 1999. Overview of hemlock health. In: McManus, K.A., K.S. Shields, and D.R. Souto eds. Proceedings: Symposium on Sustainable Management of Hemlock Ecosystems in Eastern North America, June 22-24, 1999. U.S. Forest Service, Northeastern Forest Experiment Station, General Technical Report NE-267. Durham, NH: 76-80.
  16. U.S. Forest Service (USFS). 2002. Hemlock Woolly Adelgid Management Brief. USDA Forest Service, Northeastern Area, Newtown Square, PA.