Puma concolor

(Linnaeus, 1771)

Cougar

G5Secure Found in 308 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Least concernIUCN
PSESA Status
Identity
Unique IDELEMENT_GLOBAL.2.101637
Element CodeAMAJH04010
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
CITESAppendix II
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassMammalia
OrderCarnivora
FamilyFelidae
GenusPuma
USESAPS
COSEWICPS:DD
Synonyms
Felis concolor(Linnaeus, 1771)
Other Common Names
Cougar (FR) Mountain Lion (EN) Onça-Parda, Puma, Suçuarana (PT) Puma (EN) Puma, León Americano (ES)
Concept Reference
Jones, C., R. S. Hoffman, D. W. Rice, M. D. Engstrom, R. D. Bradley, D. J. Schmidly, C. A. Jones, and R. J. Baker. 1997. Revised checklist of North American mammals north of Mexico, 1997. Occasional Papers, Museum of Texas Tech University 173:1-20.
Taxonomic Comments
Formerly included in the genus Felis.

Mitochondrial DNA analysis by Culver et al. (2000) revealed genetic uniformity across all regions of North America, relative to Central and South American populations, Culver et al. (2000) postulate that cougars were extirpated from North America during the Pleistocene extinctions of many large mammals, and only recently (circa 10,000 years ago) reinvaded North America; they suggest only one subspecies should be recognized.
Conservation Status
Rank MethodExpertise without calculation
Review Date2016-04-04
Change Date1996-11-19
Edition Date1995-05-26
Edition AuthorsGriffin, J., & G. Hammerson
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences81 to >300
Rank Reasons
Widely distributed from Canada to South America; habitat has been reduced to primarily the most remote and inaccessible areas, but the species remains relatively common (for a top predator) in several regions of the western U.S. (game species in some states) and probably elsewhere to the south of the U.S.
Range Extent Comments
Historically had widest distribution of any native American mammal (other than humans); from Canada south to southern Chile and southern Argentina and from the Pacific to the Atlantic coast. In eastern North America, now definitely known to occur only in southern Florida and Upper Peninsula of Michigan (Evers 1992). Possibly a small population exists in southeastern Canada; see Stocek (1995) for a review of recent reports from the Maritime Provinces. Elsewhere in North America, currently restricted mainly to mountainous, relatively unpopulated areas. Sea level to 14,800 ft.
Threat Impact Comments
Decline in range and abundance was due in part to federal predator control efforts augmented by state bounty programs. Loss of remote, undisturbed habitat is a problem in some areas. Excessive killing by humans may be a problem in some areas south of the U.S. Depleted ungulate populations are a potential threat.
Ecology & Habitat

Description

A large cat with an elongate body, powerful limbs, small head, short face, short rounded ears, and long neck and tail; two color phases: buff, cinnamon, and tawny to cinnamon rufous and ferruginous, and silvery gray to bluish and slaty gray; young are buffy with dark spots, and the eyes are blue for the first few months; color of upperparts is most intense midorsally; sides of muzzle and backs of ears are black; underparts are dull whitish with buff wash across the belly; end of tail is dark brown or blackish; adult total length 171-274 cm in males, 150-233 cm in females; adult tail length 53-81 cm; mass 36-120 kg in males, usually 29-64 kg in females; greatest length of skull 172-237 mm in males, 158-203 mm in females (Nowak 1991, Hall 1981, Maehr 1992, Wilson and Ruff 1999).

Diagnostic Characteristics

Differs from the jaguarundi in much larger size (maximum total length of jaguarundi is 137 cm). Differs from the lynx and bobcat in having a much longer tail (less than 25 cm in bobcat and lynx). Differs from other cats in lack of spotting in adult pelage. Young mountain lion differs from the ocelot in having the spot not arranged in rows or chainlike streaks. See Hoffmeister (1986) for cranial differences between mountain lion and jaguar.

Habitat

Now associated generally with mountainous or remote undisturbed areas. May occupy wide variety of habitats: swamps, riparian woodlands, broken country with good cover of brush or woodland. Beier (1993) determined that habitat areas of at least 2200 sq km are needed to ensure long-term population persistence; protection of corridors for immigration is highly desirable. Young are born in secluded places among rocks or dense vegetation (e.g., see Beier et al. 1995, Bleich et al. 1996).

Ecology

Primarily solitary in some areas, extensive overlap of home ranges in other areas (see Pierce et al. 1999). In Idaho, mutual avoidance maintains density of breeding adults below level set by food supply.

Annual home range varies greatly in different areas (13-1454 sq km); home range of male (generally 200 to several hundred sq km) averages larger than that of female (Kitchener 1991; Pierce et al. 1999; see also Hansen 1992 for interstate comparisons of home range size). See Beier et al. (1995) for information on movements in the Santa Ana Mountains, southern California. See Laing and Lindzey (1993, J. Mamm. 74:1056-1058) for information on replacement of individuals on vacated home ranges in Utah.

Density usually not greater than 3-4 adults per 100 sq km (8-10 per 100 sq mi) (Kitchener 1991).

Annual mortality rate in an unhunted population in Utah was 26%, over 50% in resident adults in a hunted population in Montana (see Hansen 1992).

Reproduction

Gestation lasts about 82-100 days. In the Northern Hemisphere most births occur in April-September, may occur throughout the year in Arizona. Litter size is 1-6 (usually 2-3). Young are weaned after 2-3 months. First reproduction usually occurs at 2-3 years. Young remain with mother for 1-2 years. Usually 2 years between litters (sometimes 1 year if litter does not survive). In the wild, probably few live beyond 10 years.
Terrestrial Habitats
Forest - HardwoodForest - ConiferForest - MixedWoodland - HardwoodWoodland - ConiferWoodland - MixedShrubland/chaparralDesertCliff
Palustrine Habitats
Riparian
Other Nations (2)
CanadaN4
ProvinceRankNative
OntarioSUYes
Northwest TerritoriesSUYes
SaskatchewanS2Yes
British ColumbiaS4Yes
QuebecSUYes
AlbertaS4Yes
Yukon TerritorySUYes
ManitobaSNAYes
United StatesN5
ProvinceRankNative
NevadaS5Yes
MassachusettsSXYes
LouisianaS1Yes
NebraskaS3Yes
WashingtonS4Yes
FloridaS4Yes
IdahoS5Yes
New MexicoS3Yes
UtahS4Yes
DelawareSXYes
New HampshireSHYes
IowaSXYes
District of ColumbiaSXYes
TexasS2Yes
WyomingS4Yes
MinnesotaS3Yes
MichiganS1Yes
TennesseeSXYes
KansasSXB,SNRNYes
Navajo NationS3Yes
CaliforniaSNRYes
OregonS4Yes
ArizonaS4Yes
PennsylvaniaSXYes
MontanaS5Yes
VirginiaSNRYes
Rhode IslandSXYes
ArkansasSNAYes
MarylandSHYes
KentuckySXYes
OhioSXYes
GeorgiaSHYes
North DakotaS3Yes
WisconsinSXYes
MissouriSXYes
ColoradoS4Yes
MississippiSXYes
New YorkSXYes
South DakotaS2Yes
IllinoisSXYes
MaineSHYes
VermontSHYes
ConnecticutSHYes
AlabamaSXYes
IndianaSXYes
OklahomaS1Yes
North CarolinaSXYes
New JerseySXYes
West VirginiaSHYes
South CarolinaSXYes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentHigh (continuing)
1.1 - Housing & urban areasHigh (continuing)
1.2 - Commercial & industrial areasHigh (continuing)
2 - Agriculture & aquacultureHigh (continuing)
2.1 - Annual & perennial non-timber cropsHigh (continuing)
2.2 - Wood & pulp plantationsHigh (continuing)
2.3 - Livestock farming & ranchingHigh (continuing)
4 - Transportation & service corridorsHigh (continuing)
4.1 - Roads & railroadsHigh (continuing)
5 - Biological resource useHigh (continuing)
5.1 - Hunting & collecting terrestrial animalsHigh (continuing)
5.3 - Logging & wood harvestingHigh (continuing)
6 - Human intrusions & disturbanceHigh (continuing)
6.2 - War, civil unrest & military exercisesHigh (continuing)
7 - Natural system modificationsHigh (continuing)
7.1 - Fire & fire suppressionHigh (continuing)
7.2 - Dams & water management/useHigh (continuing)
8 - Invasive & other problematic species, genes & diseasesHigh (continuing)
8.2 - Problematic native species/diseasesHigh (continuing)

Roadless Areas (308)
Arizona (20)
AreaForestAcres
Arnold MesaPrescott National Forest12,286
Black CanyonPrescott National Forest10,683
BoulderTonto National Forest40,359
Campbell BlueApache-Sitgreaves National Forests7,003
Cdo WsaCoronado National Forest1,955
CenterfireApache-Sitgreaves National Forests13,130
ChiricahuaCoronado National Forest76,876
GaliuroCoronado National Forest28,333
HackberryPrescott National Forest914
HackberryCoconino National Forest17,885
Lower San FranciscoApache-Sitgreaves National Forests59,310
Middle Dragoon RoadlessCoronado National Forest10,543
Middle Romero WSRCoronado National Forest60
NolanApache-Sitgreaves National Forests6,780
PicachoTonto National Forest4,969
SalomeTonto National Forest2,932
Santa TeresaCoronado National Forest8,929
Sierra Ancha Wilderness ContiguousTonto National Forest7,787
TumacacoriCoronado National Forest44,594
WhetstoneCoronado National Forest20,728
California (120)
AreaForestAcres
AgnewSequoia National Forest9,561
Arroyo SecoAngeles National Forest4,703
Barker ValleyCleveland National Forest11,940
Bear MountainLos Padres National Forest913
Big Butte ShinboneMendocino National Forest4,265
Black ButteLos Padres National Forest5,116
Black ButteMendocino National Forest15,461
Black CanyonInyo National Forest32,421
Black CinderLassen National Forest239
Black Mtn.Sequoia National Forest15,102
Bucks LakePlumas National Forest680
Cactus Springs BSan Bernardino National Forest3,106
CajonSan Bernardino National Forest7,548
CalienteCleveland National Forest5,953
CamuesaLos Padres National Forest8,209
Chalk PeakLos Padres National Forest7,472
ChannellSequoia National Forest45,429
ChicoSequoia National Forest39,836
Chips CreekPlumas National Forest12,940
Chips CreekLassen National Forest29,089
Circle MountainSan Bernardino National Forest6,375
City CreekSan Bernardino National Forest9,997
ColdwaterCleveland National Forest8,402
Cow CreekShasta-Trinity National Forest22,627
Coyote SoutheastInyo National Forest53,159
Cub CreekLassen National Forest8,643
Cucamonga AAngeles National Forest1,249
CuyamaLos Padres National Forest19,631
De La GuerraLos Padres National Forest5,418
Deep CreekSan Bernardino National Forest23,869
Devil GulchSierra National Forest30,490
DiableLos Padres National Forest19,597
Domeland Add.Sequoia National Forest3,046
Dry LakesLos Padres National Forest17,043
Eagle PeakCleveland National Forest6,481
East ForkShasta-Trinity National Forest6,201
Excelsior (CA)Inyo National Forest45,607
Fish CanyonAngeles National Forest29,886
Fox MountainLos Padres National Forest52,072
Garcia MountainLos Padres National Forest7,850
Glass MountainInyo National Forest52,867
Greenhorn CreekSequoia National Forest28,226
GrindstoneMendocino National Forest26,031
Grouse LakesTahoe National Forest19,085
Hixon FlatSan Bernardino National Forest8,095
Hoover - Green Ck NoHumboldt-Toiyabe National Forest7,155
Hoover - NorthHumboldt-Toiyabe National Forest1,574
Hoover - Virginia LksHumboldt-Toiyabe National Forest5,050
Horse Creek RidgeSan Bernardino National Forest8,969
HortonInyo National Forest5,717
Iceberg - Mill CreekHumboldt-Toiyabe National Forest26,988
IshiLassen National Forest21,805
JuncalLos Padres National Forest12,289
Kings RiverSierra National Forest52,999
La BreaLos Padres National Forest14,031
La PanzaLos Padres National Forest4,954
LaddCleveland National Forest5,300
LavasModoc National Forest25,864
Little French CShasta-Trinity National Forest11,529
Log Cabin SaddlebagInyo National Forest15,165
Lpoor CanyonLos Padres National Forest13,762
Machesna MountainLos Padres National Forest12,271
Magic MountainAngeles National Forest15,542
Malduce BuckhornLos Padres National Forest14,177
ManzanaLos Padres National Forest2,101
MatilijaLos Padres National Forest5,218
Middle YubaTahoe National Forest7,379
Mill CreekSequoia National Forest27,643
Mill PeakSan Bernardino National Forest7,884
MonoLos Padres National Forest28,141
MosesSequoia National Forest22,077
Mt. BidwellModoc National Forest11,687
Mt. EddyShasta-Trinity National Forest7,232
Mt. Shasta BShasta-Trinity National Forest2,809
Mt. VidaModoc National Forest7,771
No NameCleveland National Forest4,897
NordhoffLos Padres National Forest12,031
North Fork Middle Fork American RiverTahoe National Forest11,245
North Fork SmithSix Rivers National Forest37,898
PaiuteInyo National Forest58,712
PattisonShasta-Trinity National Forest29,299
Pine CreekCleveland National Forest503
Pleasant ViewAngeles National Forest26,395
PortugueseKlamath National Forest18,915
Pyramid Peak BSan Bernardino National Forest7,194
Raymond PeakStanislaus National Forest3,646
Raywood Flat BSan Bernardino National Forest11,373
Red MountainAngeles National Forest8,034
RinconSequoia National Forest54,610
Rock Creek WestInyo National Forest3,626
Rouse HillSan Bernardino National Forest13,745
RubiconEldorado National Forest4,872
San Gabriel AddAngeles National Forest2,527
San JoaquinSierra National Forest22,474
San SevaineSan Bernardino National Forest6,866
Santa CruzLos Padres National Forest21,182
Sawmill - BadlandsLos Padres National Forest51,362
Sespe - FrazierLos Padres National Forest106,910
Sespe - FrazierAngeles National Forest4,254
Sheep MountainAngeles National Forest21,098
Sill HillCleveland National Forest5,294
Slate Mtn.Sequoia National Forest12,299
Snow MountainLassen National Forest2,165
SoldierModoc National Forest10,102
South SierraSequoia National Forest8,008
South SierraInyo National Forest41,853
Strawberry PeakAngeles National Forest7,245
SugarloafSan Bernardino National Forest8,206
TequepisLos Padres National Forest9,080
ThatcherMendocino National Forest16,652
Thomes CreekMendocino National Forest16,616
Timbered CraterLassen National Forest4,096
TinemahaInyo National Forest27,060
TrabucoCleveland National Forest23,341
Trail LakeLassen National Forest1,124
WestforkAngeles National Forest4,407
Wheeler RidgeInyo National Forest15,744
White LedgeLos Padres National Forest18,632
Wilderness ContiguousMendocino National Forest3,606
WoodpeckerSequoia National Forest11,936
Colorado (7)
AreaForestAcres
Bristol HeadRio Grande NF46,087
Dome PeakRoutt NF35,716
HermosaSan Juan NF148,103
Long ParkRoutt NF42,100
Red TableWhite River NF39,122
Storm PeakSan Juan NF57,617
Troublesome SouthRoutt NF47,359
Idaho (4)
AreaForestAcres
Italian PeakCaribou-Targhee National Forest141,158
PalisadesCaribou-Targhee National Forest122,002
Pioneer MountainsSalmon-Challis National Forest172,460
Pioneer MountainsSawtooth National Forest119,563
Montana (26)
AreaForestAcres
Alexander Creek #696Kootenai National Forest6,719
Barren Creek #183Kootenai National Forest14,534
Bear - Marshall - Scapegoat - SwanLolo National Forest118,485
Bear - Marshall - Scapegoat - SwanLewis and Clark National Forest344,022
Big Snowy Mountains WsaLewis and Clark National Forest88,003
Bmss Ra 1485Flathead National Forest334,275
Buckhorn Ridge (MT)Kootenai National Forest34,716
Cabin Creek Wildlife Management Area OcdGallatin National Forest35,048
Cabinet Face East #671Kootenai National Forest50,326
Coal Ridge Ra 1127Flathead National Forest15,429
Deadhorse Ridge Ra 1128Flathead National Forest23,648
FleecerBeaverhead-Deerlodge National Forest31,585
Freezeout MountainBeaverhead-Deerlodge National Forest97,305
Gallatin FringeGallatin National Forest51,571
Hyalite - Porcupine - Buffalo Horn Wilderness Study AreaGallatin National Forest143,991
Italian PeakBeaverhead-Deerlodge National Forest90,401
Jerico MountainHelena National Forest8,445
Lazyman GulchHelena National Forest11,614
LionheadGallatin National Forest33,549
MadisonGallatin National Forest127,859
Middle Fork Judith WsaLewis and Clark National Forest81,131
North AbsarokaGallatin National Forest159,075
Petty MountainLolo National Forest16,178
Sheep Mountain - StatelineLolo National Forest37,844
Silver KingBeaverhead-Deerlodge National Forest64,289
Willard Estelle #173Kootenai National Forest3,714
Nevada (6)
AreaForestAcres
ButlerHumboldt-Toiyabe National Forest39,470
Pearl PeakHumboldt-Toiyabe National Forest71,405
Rose - GalenaHumboldt-Toiyabe National Forest3,711
Ruby - Lamoille CynHumboldt-Toiyabe National Forest32,771
Savory MountainHumboldt-Toiyabe National Forest20,570
Snake - Pole CynHumboldt-Toiyabe National Forest766
New Mexico (25)
AreaForestAcres
Alamo CanyonSanta Fe National Forest8,639
Arroyo de los FrijolesSanta Fe National Forest5,277
Aspen MountainGila National Forest23,784
Black CanyonSanta Fe National Forest1,922
Brushy MountainGila National Forest7,199
Candian RiverCibola National Forest7,149
Columbine - Hondo Wilderness Study AreaCarson National Forest43,739
Contiguous To Black & Aldo Leopold WildernessGila National Forest111,883
Contiguous To Gila Wilderness & Primitive AreaGila National Forest79,049
Devils CreekGila National Forest89,916
GallinasSanta Fe National Forest13,208
Gila BoxGila National Forest23,759
Guaje CanyonSanta Fe National Forest6,104
Juan de Gabaldon GrantSanta Fe National Forest8,023
Latir PeakCarson National Forest3,573
LemitasSanta Fe National Forest8,129
Meadow CreekGila National Forest34,167
Nichols ReservoirSanta Fe National Forest1,518
NolanGila National Forest13,051
Pacheco CanyonSanta Fe National Forest1,012
Peloncillo (NM)Coronado National Forest43,339
Pueblo MesaSanta Fe National Forest3,540
RendijaSanta Fe National Forest2,176
Ryan HillCibola National Forest34,201
South Guadalupe MountainsLincoln National Forest20,930
Oregon (22)
AreaForestAcres
BuckhornWallowa-Whitman National Forest17,180
Bull Of The WoodsMt. Hood National Forest8,843
Calf - Copeland CreekUmpqua National Forest15,696
Crane MountainFremont National Forest23,096
FairviewUmpqua National Forest7,417
French Pete (a)Willamette National Forest1,668
LakeMt. Hood National Forest1,327
Limpy RockUmpqua National Forest6,782
Lookout MountainOchoco National Forest14,115
Metolius BreaksDeschutes National Forest11,141
Middle SantiamWillamette National Forest7,316
Mt. JeffersonDeschutes National Forest2,282
North KalmiopsisSiskiyou National Forests91,560
North Mt. EmilyUmatilla National Forest4,416
OlallieMt. Hood National Forest7,607
Opal CreekWillamette National Forest5,417
PotamusUmatilla National Forest5,389
Snake RiverWallowa-Whitman National Forest31,229
TahkenitchSiuslaw National Forest5,799
TenmileSiuslaw National Forest10,818
West - South BachelorDeschutes National Forest25,994
WoahinkSiuslaw National Forest5,309
Utah (30)
AreaForestAcres
0401002Ashley National Forest36,113
0401011Ashley National Forest30,062
0401012Ashley National Forest46,400
418001Uinta National Forest5,697
418012Uinta National Forest25,758
418013Uinta National Forest14,643
418021Uinta National Forest6,255
418025Uinta National Forest32,698
418027Uinta National Forest13,884
418028Uinta National Forest34,002
418029Uinta National Forest15,673
Boulder Mtn. / Boulder Top / Deer LakeDixie National Forest110,690
Capital ReefDixie National Forest763
Casto BluffDixie National Forest87,466
Circleville MountainFishlake National Forest24,142
FishhookDixie National Forest12,959
HogsbackWasatch-Cache National Forest7,936
LakesWasatch-Cache National Forest121,967
Lone Peak ContiguousWasatch-Cache National Forest874
Long Neck Mesa / Steep Creek / Oak Creek - Steep Creek / OakDixie National Forest55,489
Mt. AireWasatch-Cache National Forest9,681
Mt. Logan SouthWasatch-Cache National Forest17,014
Mt. Logan WestWasatch-Cache National Forest5,285
NoblettsWasatch-Cache National Forest3,116
North FrancisWasatch-Cache National Forest8,148
Pine Valley MountainsDixie National Forest57,673
Right Hand Fork LoganWasatch-Cache National Forest15,023
Table Cliffs - Henderson CanyonDixie National Forest19,581
Temple PeakWasatch-Cache National Forest24,081
Upper South ForkWasatch-Cache National Forest16,811
Washington (33)
AreaForestAcres
Abercrombie - HooknoseColville National Forest33,862
Asotin CreekUmatilla National Forest16,433
Big Lava BedGifford Pinchot National Forest19,043
Black CanyonOkanogan National Forest9,681
ChelanWenatchee National Forest74,650
Glacier Peak IMt Baker-Snoqualmie National Forest15,175
Glacier Peak KMt Baker-Snoqualmie National Forest47,269
Gotchen CreekGifford Pinchot National Forest7,518
Granite MountainOkanogan National Forest27,428
Grassy TopColville National Forest10,302
Grassy TopIdaho Panhandle National Forests13,485
Green MountainOlympic National Forest4,617
Higgins MountainMt Baker-Snoqualmie National Forest13,185
Jupiter RidgeOlympic National Forest10,148
Long SwampOkanogan National Forest66,344
Madison CreekOlympic National Forest1,223
Mill Creek Watershed (WA)Umatilla National Forest16,747
Mt. Baker MaMt Baker-Snoqualmie National Forest24,847
Mt. Baker Noisy - DiobsudMt Baker-Snoqualmie National Forest56,039
Mt. Baker WestMt Baker-Snoqualmie National Forest25,390
Mt. BonaparteOkanogan National Forest10,891
Norse PeakWenatchee National Forest10,169
Pasayten RimOkanogan National Forest17,074
QuilceneOlympic National Forest18,656
Red MountainGifford Pinchot National Forest2,992
Salmo - Priest BColville National Forest11,869
SawtoothOkanogan National Forest122,194
TaneumWenatchee National Forest26,140
TeanawayWenatchee National Forest72,849
Thorp Mtn.Wenatchee National Forest22,717
TiffanyOkanogan National Forest22,045
Upper SkokomishOlympic National Forest9,311
Upper TucannonUmatilla National Forest12,485
Wyoming (15)
AreaForestAcres
Gros Ventre MountainsBridger-Teton National Forest106,418
Lake Alice - Commissary RidgeBridger-Teton National Forest166,707
Little BighornBighorn National Forest133,949
Middle ForkMedicine Bow-Routt National Forest13,238
Nugent Park - Hams Fork RidgeBridger-Teton National Forest21,241
Pacific Creek - Blackrock CreekBridger-Teton National Forest24,658
PalisadesTarghee National Forest1,121
Phillips RidgeBridger-Teton National Forest10,108
Sheep MountainMedicine Bow-Routt National Forest17,626
South ForkShoshone National Forest64,903
Spread Creek - Gros Ventre RiverBridger-Teton National Forest166,097
Walker PrairieBighorn National Forest62,434
Wapiti Valley NorthShoshone National Forest18,345
Wapiti Valley SouthShoshone National Forest43,517
Wilderness Study AreaTarghee National Forest51,961
References (61)
  1. Albuja, L. 1991. Lista de vertebrados del Ecuador. Escuela Politécnica XVI:163-203.
  2. American Society of Mammalogists (ASM). 2025. Mammal Diversity Database (Version 1.13) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.10595931. Online. Available: https://www.mammaldiversity.org/
  3. Anderson, C. R., F. G. Lindzey, and D. B. McDonald. 2004. Genetic structure of cougar populations across the Wyoming Basin: metapopulation or megapopulation. Journal of Mammalogy 85:1207-1214.
  4. Banfield, A. W. F. 1974. The mammals of Canada. University of Toronto Press, Toronto, Canada. 438 pp.
  5. Barnes, C.T. 1960. The cougar or mountain lion. Salt Lake City, UT. 176 pp.
  6. Beck, W.H. 1958. A guide to Saskatchewan mammals. Special Publication No. 1. Saskatchewan Natural History Society, Regina, Saskatchewan.
  7. Beier, P. 1991. Cougar attacks on humans in the United States and Canada. Wildl. Soc. Bull. 19:
  8. Beier, P. 1993. Determining minimum habitat areas and habitat corridors for cougars. Conservation Biology 7:94-
  9. Beier, P., D. Choate, and R. H. Barrett. 1995. Movement patterns of mountain lions during different behaviors. Journal of Mammalogy 76:1056-1070.
  10. Belden, R. C., et al. 1988. Panther habitat use in southern Florida. J. Wildlife Management 52:660-663.
  11. Bleich, V. C., B. M. Pierce, J. L. Davis, and V. L. Davis. 1996. Thermal characteristics of mountain lion dens. Great Basin Naturalist 56:276-278.
  12. Bradley, R.D., L.K. Ammerman, R.J. Baker, L.C. Bradley, J.A. Cook. R.C. Dowler, C. Jones, D.J. Schmidly, F.B. Stangl Jr., R.A. Van den Bussche and B. Würsig. 2014. Revised checklist of North American mammals north of Mexico, 2014. Museum of Texas Tech University Occasional Papers 327:1-28. Available at: http://www.nsrl.ttu.edu/publications/opapers/ops/OP327.pdf
  13. Cadena, A., R. P. Anderson, and P. Rivas-Pava. 1998. Colombian mammals from the chocoan slopes of Nariño. Occasional Papers Museum of Texas Tech University 180:1-15.
  14. Caire, W., J. D. Tyler, B. P. Glass, and M. A. Mares. 1989. Mammals of Oklahoma. University of Oklahoma Press, Norman. Oklahoma. 567 pp.
  15. Carrillo, E., G. Wong, and A. Cuarón. 2000. Monitoring mammal populations in Costa Rican protected areas under different hunting restrictions. Conservation Biology 14:1580-1591.
  16. Culver, M., W. E. Johnson, J. Pecon-Slattery, and S. J. O'Brien. 2000. Genomic ancestry of the American puma (<i>Puma concolor</i>). Journal of Heredity 91:186-197.
  17. Currier, M.J.P. 1983. FELIS CONCOLOR. Mammalian species, 200:1-7.
  18. Dixon, K. R. 1982. Mountain lion. Pages 711-727 in Chapman, J. A., and G. A. Feldhammer, editors. Wild mammalsof North America. Biology, management, and economics. Johns Hopkins Univ. Press, Baltimore.
  19. Eaton, R.L. 1973. The status, management, and conservation of the cougar in the United States. The World's Cats, 1:68-86.
  20. Eisenberg, J. F. and K. H. Redford. 1989. Mammals of the Neotropics, The Northern Neotropics, Volume 1, Panama, Colombia, Venezuela, Guyana, Suriname, French Guiana. Chicago, The University of Chicago Press.
  21. Eisenberg, J. F. and K. H. Redford. 1999. Mammals of the Neotropics, The Central Neotropics, Volume 3, Ecuador, Peru, Bolivia, Brazil. Chicago, The University of Chicago Press.
  22. Emmons, L. H. and F. Feer. 1997. Neotropical rainforest mammals, a field guide. Second edition. Chicago, The University of Chicago Press.
  23. Engstrom, M. and B. Lim. 2000. Checklist of the mammals of Guyana. Smithsonian Institute. Http://www.mnh.si.edu/biodiversity/bdg/guymammals.html
  24. Evers, D. C. 1992. A guide to Michigan's endangered wildlife. Univ. Michigan Press, Ann Arbor. viii + 103 pp.
  25. Hall, E. R. 1981a. The Mammals of North America, second edition. Vols. I &amp; II. John Wiley &amp; Sons, New York, New York. 1181 pp.
  26. Handley, C. O., Jr. 1991. Mammals. Pages 539-616 in K. Terwilliger, coordinator. Virginia's endangered species: proceedings of a symposium. McDonald and Woodward Publishing Company, Blacksburg, Virginia.
  27. Hansen, K. 1992. Cougar: the American lion. Northland Publising Company, Flagstaff, Arizona. xiii + 129 pp.
  28. Hebda, A.J. 2011. List of mammals of Nova Scotia (including synonyms used in the literature relating to Nova Scotia) (revision 2) 24 July 2011. Nova Scotia Museum Collections Unit, Halifax, Nova Scotia. 24 pp. Online. Available: https://naturalhistory.novascotia.ca/sites/default/files/inline/images/names_and_synonyms_ver3.pdf
  29. Hoffmeister, D. F. 1986. Mammals of Arizona. University of Arizona Press and Arizona Game and Fish Department. 602 pp.
  30. Jones, C., R. S. Hoffman, D. W. Rice, M. D. Engstrom, R. D. Bradley, D. J. Schmidly, C. A. Jones, and R. J. Baker. 1997. Revised checklist of North American mammals north of Mexico, 1997. Occasional Papers, Museum of Texas Tech University 173:1-20.
  31. Jones, J. K., Jr., D. M. Armstrong, R. S. Hoffmann, and C. Jones. 1983. Mammals of the Northern Great Plains. University of Nebraska Press, Lincoln, Nebraska.
  32. Jones, J. K., Jr., R. S. Hoffman, D. W. Rice, C. Jones, R. J. Baker, and M. D. Engstrom. 1992a. Revised checklist of North American mammals north of Mexico, 1991. Occasional Papers, The Museum, Texas Tech University, 146:1-23.
  33. Kitchener, A. 1991. The natural history of the wild cats. Comstock Publishing Associates, Cornell Univ. Press, Ithaca. xxi + 280 pp.
  34. Lindzey, F. 1987. Mountain Lion. Pp. 656-668 in: Novak, M., J.A. Baker, M.E. Obbard &amp; B. MALLOCH (EDS.). 1987. Wild furbearer management and conservation in North America. Ontario Ministry of Natural Resources. Toronto. 1150 pp.
  35. Linzey, D.W. 2016. Mammals of Great Smoky Mountains National Park: 2016 revision. Southeastern Naturalist 15(Monograph 8):1–93.
  36. Logan, K. A., and L. L. Sweanor. 2001. Desert puma: evolutionary ecology and conservation of an enduring carnivore. Island Press, Covelo, California. 463 pp.
  37. Maehr, D. S., and J. B. Wooding. 1992. Florida black bear <i>Ursus americanus floridanus</i>. Pages 265-275 in S. R. Humphrey, editor. Rare and endangered biota of Florida. Vol. I. Mammals. Univ. Press of Florida, Gainesville. xviii + 392 pp.
  38. Matthews, J.R. and C.J. Moseley (eds.). 1990. The Official World Wildlife Fund Guide to Endangered Species of North America. Volume 1. Plants, Mammals. xxiii + pp 1-560 + 33 pp. appendix + 6 pp. glossary + 16 pp. index. Volume 2. Birds, Reptiles, Amphibians, Fishes, Mussels, Crustaceans, Snails, Insects, and Arachnids. xiii + pp. 561-1180. Beacham Publications, Inc., Washington, D.C.
  39. Moorhead, B., and T. Hofstra. 1994. Western park personnel meet on mountain lion-human encounters. Park Science, Fall 1994, pp. 20-21.
  40. Nowak, R. M. 1991. Walker's mammals of the world. Fifth edition. Vols. I and II. Johns Hopkins Univ. Press, Baltimore. 1629 pp.
  41. Nowell, K., and P. Jackson. 1996. Wild cats: Status survey and conservation action plan. Gland, Switzerland, IUCN.
  42. Pacheco, V., H. de Macedo, E. Vivar, C. Ascorra, R. Arana-Cardó, and S. Solari. 1995. Lista anotada de los mamíferos peruanos. Conservation International, Washington, DC.
  43. Pacheco, V., S. Solari, E. Vivar, and P. J. Hocking. 1994. La riqueza biológica del Parque Nacional Yanachaga - Chemillén. Magistri et Doctores 7:3-6.
  44. Pierce, B. M., V. C. Bleich, J. D. Wehausen, and R. T. Bowyer. 1999. Migratory patterns of mountain lions: implications for social regulation and conservation. Journal of Mammalogy 80:986-992.
  45. Ross, I. P., and M. G. Jalkotzy. 1995. Fates of translocated cougars, FELIS CONCOLOR, in Alberta. Canadian Field-Naturalist 109:475-476.
  46. Seal, U. S., et al. 1989. Florida panther FELIS CONCOLOR CORYI viability analysis and species survival plan. Captive Breeding Specialist Group, IUCN.
  47. Seidensticker, J. C., IV, M. C. Hornocker, W. V. Wiles, and J. P. Messick. 1973. Mountain lion social organization in the Idaho Primitive Area. Wildlife Monogr. 35:1-60.
  48. Stocek, R. F. 1995. The cougar, FELIS CONCOLOR, in the Maritime Provinces. Canadian Field-Naturalist 109:19-22.
  49. Sweanor, L. L., K. A. Logan, and M. G. Hornocker. 2000. Cougar dispersal patterns, metapopulation dynamics, and conservation. Conservation Biology 14:798-808.
  50. Tirira, D. 1999. Mamíferos del Ecuador. Museo de Zoología, Pontificia Universidad Católica del Ecuador, Quito.
  51. Tischendorf, J. W., and S. J. Ropski, editors. 1996. Proceedings of the eastern cougar conference, 1994. American Ecological Research Institute. 245 pp.
  52. U.S. Fish and Wildlife Service (USFWS). 1980. Selected vertebrate endangered species of the seacoast of the United States- the eastern cougar. FWS/OBS-80/01.44, Slidell.
  53. Van Dyke, F. G., et al. 1986. Reactions of mountain lions to logging and human activity. J. Wildl. Manage. 50:95-102.
  54. Van Sickle, W. D., and F. G. Lindzey. 1991. Evaluation of a cougar population estimator based on probability sampling. J. Wildl. Manage. 55:738-743.
  55. Van Sickle, W. D., and F. G. Lindzey. 1992. Evaluation of road track surveys for cougars (FELUS CONCOLOR). Great Basin Nat. 52:232-236.
  56. White, T. 1982. Saskatchewan Cougar - Elusive Cat. Saskatchewan Natural History Society, Special Publication. 80 pp.
  57. Wilson, D. E., and D. M. Reeder (editors). 1993. Mammal species of the world: a taxonomic and geographic reference. Second edition. Smithsonian Institution Press, Washington, DC. xviii + 1206 pp. Available online at: http://www.nmnh.si.edu/msw/.
  58. Wilson, D. E., and D. M. Reeder (editors). 2005. Mammal species of the world: a taxonomic and geographic reference. Third edition. The Johns Hopkins University Press, Baltimore. Two volumes. 2,142 pp. [As modified by ASM the Mammal Diversity Database (MDD) at https://www.mammaldiversity.org/index.html]
  59. Wilson, D. E., and S. Ruff. 1999. The Smithsonian book of North American mammals. Smithsonian Institution Press, Washington, D.C. 750 pp.
  60. Yanez, J. L., et al. 1986. Food habits of the southernmostmountain lions (FELIS CONCOLOR) in South America: natural versus livestocked ranges. J. Mamm. 67:604-606.
  61. Young, S. P. and E. A. Goldman. 1946. The Puma, mysterious American cat. Part I (by Young). History, life habits, economic status, and control. Part II (by Goldman). Classification of the races of the puma. American Wildlife Institute (also Dover Publ., Inc., New York). 358 PP.