Plethodon richmondi

Netting and Mittleman, 1938

Southern Ravine Salamander

G5Secure Found in 15 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Least concernIUCN
LowThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.103596
Element CodeAAAAD12150
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicendemic to a single nation
KingdomAnimalia
PhylumCraniata
ClassAmphibia
OrderCaudata
FamilyPlethodontidae
GenusPlethodon
Concept Reference
Frost, D. R. 1985. Amphibian species of the world. A taxonomic and geographical reference. Allen Press, Inc., and The Association of Systematics Collections, Lawrence, Kansas. v + 732 pp.
Taxonomic Comments
This salamander was included in P. cinereus until 1938, when it was described as P. richmondi.

Plethodon electromorphus formerly was included in this species (see Highton 1999). Highton (1999) described Plethodon electromorphus as distinct from Plethodon richmondi on the basis of protein characteristics detectable by eletrophoresis; P. electromorphus and P. richmondi do not share common alleles at one of 24 genetic loci and have a fixed difference at another locus in all but one population (Highton 1999). These two salamanders hybridize in a narrow zone of contact, with low introgression (Highton 1999). Plethodon richmondi and P. cinereus hybridize on Iron and Holston Mountain, Tennessee (Thurow 1969). Some taxonomists would regard P. electromorphus as questionably valid as a species distinct from P. richmondi.
Conservation Status
Review Date2002-03-26
Change Date2001-11-09
Edition Date2011-05-06
Edition AuthorsHammerson, G.
Threat ImpactLow
Range Extent20,000-200,000 square km (about 8000-80,000 square miles)
Number of Occurrences81 - 300
Rank Reasons
Relatively common and widely distributed in eastern Kentucky, southern West Virginia, western Virginia, northwestern North Carolina, and northeastern Tennessee.
Range Extent Comments
Range includes eastern Kentucky, southern West Virginia (west of the New and Kanawha rivers and south of the Teays River valley), western Virginia, northwestern North Carolina, and northeastern Tennessee (Highton 1999).
Occurrences Comments
This species is represented by many and/or large occurrences (subpopulations) throughout most of the range (Regester 2000).
Threat Impact Comments
Deforestation and urbanization have eliminated local populations (Petranka 1998).
Ecology & Habitat

Habitat

Terrestrial. Found under logs, stumps, rocks, or leaf litter on moist wooded slopes of valleys and ravines; high densities on wooded talus; rarely on dry ridge crests, hilltops, or valley floors (Regester 2000). Generally underground in mid-winter and summer. Lays eggs under rocks or in underground cavity.

Reproduction

Lays cluster of about 6 eggs in spring. Larval stage passed in egg. Hatching occurs in August-September.
Terrestrial Habitats
Forest - HardwoodWoodland - Hardwood
Palustrine Habitats
Riparian
Other Nations (1)
United StatesN5
ProvinceRankNative
OhioSNRYes
KentuckyS5Yes
North CarolinaS3Yes
VirginiaS4Yes
West VirginiaS4Yes
TennesseeS3Yes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentHigh (continuing)
1.1 - Housing & urban areasHigh (continuing)
5 - Biological resource useHigh (continuing)
5.3 - Logging & wood harvestingHigh (continuing)

Roadless Areas (15)
Kentucky (1)
AreaForestAcres
WolfpenDaniel Boone National Forest2,835
North Carolina (1)
AreaForestAcres
Wilson CreekPisgah National Forest4,863
Tennessee (4)
AreaForestAcres
Big Laurel Branch AdditionCherokee National Forest5,577
Flint Mill GapCherokee National Forest9,494
London Bridge BranchCherokee National Forest3,387
Rogers RidgeCherokee National Forest4,738
Virginia (9)
AreaForestAcres
Bear CreekJefferson National Forest18,274
Garden MountainJefferson National Forest3,960
Hunting Camp Little Wolf CreekJefferson National Forest8,953
Lewis Fork AdditionJefferson National Forest749
Little Wilson Creek Addition BJefferson National Forest1,725
New London Bridge BranchJefferson National Forest844
North Fork PoundJefferson National Forest4,757
Raccoon BranchJefferson National Forest4,388
Seng MountainJefferson National Forest6,428
References (13)
  1. Barbour, R. W. 1971. Amphibians and reptiles of Kentucky. Univ. Press of Kentucky, Lexington. x + 334 pp.
  2. Behler, J. L., and F. W. King. 1979. The Audubon Society field guide to North American reptiles and amphibians. Alfred A. Knopf, New York. 719 pp.
  3. Blackburn, L., P. Nanjappa, and M. J. Lannoo. 2001. An Atlas of the Distribution of U.S. Amphibians. Copyright, Ball State University, Muncie, Indiana, USA.
  4. Crother, B. I. (editor). 2017. Scientific and standard English names of amphibians and reptiles of North America north of Mexico, with comments regarding confidence in our understanding. 8th edition. SSAR Herpetological Circular 43:1-104. [Updates in SSAR North American Species Names Database at: https://ssarherps.org/cndb]
  5. Frost, D. R. 1985. Amphibian species of the world. A taxonomic and geographical reference. Allen Press, Inc., and The Association of Systematics Collections, Lawrence, Kansas. v + 732 pp.
  6. Frost, D.R. 2020. Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History, New York, USA. Online: http://research.amnh.org/herpetology/amphibia/index.html
  7. Green, N. B., and T. K. Pauley. 1987. Amphibians and reptiles in West Virginia. University of Pittsburgh Press, Pittsburgh, Pennsylvania. xi + 241 pp.
  8. Highton, R. 1999a. Geographic protein variation and speciation in the salamanders of the <i>Plethodon cinereus</i> group with the description of two new species. Herpetologica 55:43-90.
  9. Highton, R. 1999b. Hybridization in the contact zone between <i>Plethodon richmondi</i> and <i>Plethodon electromorphus</i> in northern Kentucky. Herpetologica 55:91-105.
  10. Mahoney, M. J. 2001. Molecular systematics of <i>Plethodon </i>and <i>Aneides </i>(Caudata: Plethodontini): phylogenetic analysis of an old and rapid radiation. Molecular Phylogenetics and Evolution 18:174-188.
  11. Minton, S. A., Jr. 1972. Amphibians and reptiles of Indiana. Indiana Academy Science Monographs 3. v + 346 pp.
  12. Petranka, J. W. 1998. Salamanders of the United States and Canada. Smithsonian Institution Press, Washington, D.C.
  13. Regester, K.J. 2000. Plethodon richmondi. Catalogue of American Amphibians and Reptiles. 707:1-3.