Dryophytes chrysoscelis

(Cope, 1880)

Cope's Gray Treefrog

G5Secure Found in 23 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Least concernIUCN
Identity
Unique IDELEMENT_GLOBAL.2.103778
Element CodeAAABC02050
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassAmphibia
OrderAnura
FamilyHylidae
GenusDryophytes
Synonyms
Hyla chrysoscelisCope, 1880
Other Common Names
Cope's gray treefrog (EN) Cope's Grey Treefrog (EN) Rainette criarde (FR)
Concept Reference
Frost, D. R. 1985. Amphibian species of the world. A taxonomic and geographical reference. Allen Press, Inc., and The Association of Systematics Collections, Lawrence, Kansas. v + 732 pp.
Taxonomic Comments
Duellman et al. (2016) restricted Hyla to Eurasia and North Africa and referred the North American and east Asian sister taxon of this group to Dryophytes. This taxonomy was accepted by Amphibian Species of the World and followed here. It is not followed by the Society for the Study of Amphibians And Reptiles (SSAR) with Crother (2017) stating, "acceptance of this taxonomy within the community is not clear at this point."

Faivovich et al. (2005) redelimited this monophyletic taxon to include only North American and Eurasian species. Hua et al. (2009) discussed relationships within the group. Fouquette and Dubois (2014) recognized a suite of subgenera based on genetic and morphological evidence, but pending a more thorough evidentiary review, Crother (2017) hesitates to employ this taxonomy.

Not distinguished from look-alike species H. versicolor in much published literature; distinguished by chromosomes, erythrocyte size (Matson 1990), and call characteristics. Natural hybridization between H. versicolor and H. chrysoscelis has been confirmed, but apparently there is severe selection against hybrids (sexually mature hybrids are very rare) (Gerhardt et al. 1994).

Two chromosome morphs reported in chrysoscelis (Wiley 1983); see also Ralin et al. (1983).
Conservation Status
Rank MethodExpertise without calculation
Review Date2015-06-05
Change Date2001-10-26
Edition Date2010-01-26
Edition AuthorsHammerson, G.
Rank Reasons
Large range in much of eastern United States and a small part of south-central Canada; many secure occurrences.
Range Extent Comments
Range is not precisely known but includes most of the south-central and southeastern United States and areas west of the Great Lakes, from Manitoba, Minnesota, Wisconsin, Michigan, southern Ohio, West Virginia, and Maryland south to eastern Texas, the Gulf Coast, and northern Florida (Preston 1982, Holloway et al. 2006). See Little et al. (1989) for information on distribution in West Virginia, southern Ohio, and southwestern Pennsylvania.
Occurrences Comments
Represented by many and/or large occurrences throughout most of the range.
Ecology & Habitat

Description

This species is essentially identical to the gray treefrog (Hyla versicolor) The upper side has numerous small warts and is usually green (especially juveniles) to gray and often has a pattern that resembles lichens that grow on tree trunks. There is a light spot under each eye. The groin and concealed bases of the hind legs are orange-yellow with black mottling. Maximum size is about 2.5 inches (6 cm) snout-vent length. Breeding male can be recognized by their darl loose throat skin. Breeding calls are loud trills (often mistaken for a woodpecker's call), with pulses emitted faster than in the gray treefrog (Hyla versicolor). Larvae have strongly arched tails fins that may be heavily mottled with black and often tinged red or orange, and (if not broken) ending in a slender filament. Larvae reach a total length of up to around 1.5 inches (3.8 cm). Egg masses contain clusters of about 6-45 eggs, floating free or loosely attached to submerged vegetation.

Habitat

Cope's gray treefrogs inhabit wooded areas and woodland edges (including woodlots in prairies), usually within a few hundred meters of the aquatic habitats in whch they breed. Often they occur in recently disturbed areas with abundant shrubs, herbaceous growth, and vines. Activity is arboreal and terrestrial. In Tennessee, frogs associated with knothole cavities in trees in fall were not there after mid-November (Ritke and Babb 1991). When inactive, frogs may hide in tree holes, under bark, under leaves, or under tree roots.

Breeding sites include temporary or permanent waters of flooded ditches, puddles, river sloughs, creeks, and small ponds, where there are woody branches or extensive herbaceous growth along the edges. Males call from the water surface or from vegetation or ground near water. Individuals generally breed in the same site in successive years (Ritke et al. 1991).

Ecology

In Tennessee, the majority of the adults marked in one year were not recaptured the following year, probably due to predation and/or winter mortality (or possibly postponed reproduction) (Ritke et al. 1991).

Skin secretions of these frogs are noxious to certain potential predators and may cause irritation if they contact human eyes or other sensitive membranes.

Reproduction

Breeding occurs in spring-early summer (May-early July in Kansas, April-July in Maryland, April-August in western Tennessee). Often, but not always, breeding is stimulated by rainfall and warm temperatures. A female's clutch is divided among small clusters of around 20-40 eggs, deposited at the water's surface or attached to emergent plants. Individual females produce 1-3 clutches/year in Kansas and Tennessee (most females produce one clutch/year). Eggs hatch in several days. Larvae metamorphose in about 6-9 weeks, by end of summer (may rarely overwinter; McCallum and McCallum, 2005, Herpetol. Rev. 36:54). Males mated up to 3 times per season in Kansas; most calling males did not mate (Godwin and Roble 1983, Ritke et al. 1990). Females in Tennessee apparently require at least two years to reach sexual maturity.
Terrestrial Habitats
Forest - HardwoodForest - MixedWoodland - HardwoodWoodland - MixedShrubland/chaparral
Palustrine Habitats
TEMPORARY POOLHERBACEOUS WETLANDSCRUB-SHRUB WETLANDFORESTED WETLANDRiparian
Other Nations (2)
CanadaN4
ProvinceRankNative
ManitobaS4Yes
United StatesN5
ProvinceRankNative
North CarolinaS5Yes
MissouriS5Yes
VirginiaS5Yes
ArkansasS4Yes
TennesseeS5Yes
LouisianaS5Yes
MarylandS5Yes
MississippiS5Yes
West VirginiaS4Yes
FloridaS4Yes
GeorgiaS5Yes
New JerseyS2Yes
DelawareS2Yes
OklahomaSNRYes
NebraskaS5Yes
PennsylvaniaS1Yes
South CarolinaS5Yes
TexasS5Yes
South DakotaS2Yes
District of ColumbiaS4Yes
AlabamaS5Yes
KansasS5Yes
WisconsinS5Yes
IowaS4Yes
MinnesotaS5Yes
MichiganS5Yes
IllinoisS4Yes
OhioSNRYes
IndianaS4Yes
KentuckyS5Yes
Roadless Areas (23)
Arkansas (1)
AreaForestAcres
Bear MountainOuachita National Forest1,910
Florida (1)
AreaForestAcres
Gum BayApalachicola National Forest11,645
Georgia (1)
AreaForestAcres
Joe GapChattahoochee National Forest5,321
North Carolina (14)
AreaForestAcres
Bald MountainPisgah National Forest11,085
BearwallowPisgah National Forest4,113
Big Indian (addition)Nantahala National Forest1,155
Cheoah BaldNantahala National Forest7,795
Craggy MountainPisgah National Forest2,657
Dobson KnobPisgah National Forest6,111
Graveyard Ridge (addition)Pisgah National Forest1,958
Harper CreekPisgah National Forest7,325
Lost CovePisgah National Forest5,944
Mackey MountainPisgah National Forest5,934
Sam Knob (addition)Pisgah National Forest2,576
South Mills RiverPisgah National Forest8,588
Wilson CreekPisgah National Forest4,863
Woods MountainPisgah National Forest9,602
Tennessee (4)
AreaForestAcres
Bald MountainCherokee National Forest11,743
Big Laurel Branch AdditionCherokee National Forest5,577
Sampson Mountain AdditionCherokee National Forest3,064
Stone MountainCherokee National Forest5,367
Virginia (2)
AreaForestAcres
New London Bridge BranchJefferson National Forest844
Seng MountainJefferson National Forest6,428
References (25)
  1. Blackburn, L., P. Nanjappa, and M. J. Lannoo. 2001. An Atlas of the Distribution of U.S. Amphibians. Copyright, Ball State University, Muncie, Indiana, USA.
  2. Collins, J. T. 1982. Amphibians and reptiles in Kansas. Second edition. Univ. Kansas Mus. Nat. Hist., Pub. Ed. Ser. 8. xiii + 356 pp.
  3. Conant, R., and J. T. Collins. 1998. A field guide to reptiles and amphibians: eastern and central North America. Third edition, expanded. Houghton Mifflin Co., Boston, Massachusetts. 616 pp.
  4. Crother, B. I. (editor). 2017. Scientific and standard English names of amphibians and reptiles of North America north of Mexico, with comments regarding confidence in our understanding. 8th edition. SSAR Herpetological Circular 43:1-104. [Updates in SSAR North American Species Names Database at: https://ssarherps.org/cndb]
  5. Duellman, W. E., A. B. Marion, and S. B. Hedges. 2016. Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae). Zootaxa 4104: 1–109.
  6. Faivovich, J., C.F.B. Haddad, P.C.A. Garcia, D.R. Frost, J.A. Campbell and W.C. Wheeler. 2005. Systematic review of the frog family Hylidae, with special reference to Hylinae: Phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of Natural History 294:1-240.
  7. Fouquette Jr., M.J., and A. DuBois. 2014. A Checklist of North American Amphibians and Reptiles. Seventh Edition. Volume 1—Amphibians. Xlibris LLC, Bloomington, Indiana. 586 pp.
  8. Frost, D. R. 1985. Amphibian species of the world. A taxonomic and geographical reference. Allen Press, Inc., and The Association of Systematics Collections, Lawrence, Kansas. v + 732 pp.
  9. Frost, D.R. 2020. Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History, New York, USA. Online: http://research.amnh.org/herpetology/amphibia/index.html
  10. Gerhardt, H.C., Ptacek, M.B., Barnett, L. and Torke, K.G 1994. Hybridization in the diploid-tetraploid treefrogs <i>Hyla chrysoscelis</i> and <i>Hyla versicolor</i>. Copeia 1994:51-59.
  11. Godwin, G. J., and S. M. Roble. 1983. Mating success in male treefrogs, HYLA CHRYSOSCELIS (Anura: Hylidae). Herpetologica 39:141-146.
  12. Holloway, A. K., D. C. Cannatella, H. C. Gerhardt, and D. M. Hillis. 2006. Polyploids with different origins and ancestors form a single sexual polyploid species. American Naturalist 167(4):E88-E101.
  13. Hua, X., C.-z. Fu, J.-t. Li, A. Nieto-Montes de Oca, and J. J. Wiens. 2009. A revised phylogeny of Holarctic treefrogs (genus <i>Hyla</i>) based on nuclear and mitochondrial DNA sequences. Herpetologica 65:246-259.
  14. Jaslow, Alan P. and Richard C. Vogt. 1977. Identification and distribution of Hyla versicolor and Hyla chrysoscelis in Wisconsin. Herpetologica 33(2):201-205.
  15. Johnson, T.R. 1977. The Amphibians of Missouri. University of Kansas Museum of Natural History, Public Education Series 6: ix + 134 pp.
  16. Little, M. A., B. L. Monroe, Jr., and J. E. Wiley. 1989. The distribution of the HYLA VERSICOLOR complex in the northern Appalachian highlands. J. Herpetol. 23:299-303.
  17. Matson, T. O. 1990. Erythrocyte size as a taxonomic character in the identification of Ohio HYLA CHRYSOSCELIS and H. VERSICOLOR. Herpetologica 46:457-462.
  18. Mount, R. H. 1975. The reptiles and amphibians of Alabama. Auburn University Agricultural Experiment Station, Auburn, Alabama. vii + 347 pp.
  19. Preston, W. B. 1982. The amphibians and reptiles of Manitoba. Manitoba Museum of Man and Nature, Winnipeg, Manitoba. 128 pp.
  20. Ralin, D. B., M. A. Romano, and C. W. Kilpatrick. 1983. The tetraploid treefrog HYLA VERSICOLOR: evidence for a single origin from the diploid H. CHRYSOSCELIS. Herpetologica 39:212-225.
  21. Ritke, M. E., and J. G. Babb. 1991. Behavior of the gray treefrog (HYLA CHRYSOSCELIS) during the non-breeding season. Herpetol. Rev. 22:5-6, 8.
  22. Ritke, M. E., J. G. Babb, and M. K. Ritke. 1990. Life history of the gray treefrog (HYLA CHRYSOSCELIS) in wetsern Tennessee. J. Herpetol. 24:135-141.
  23. Ritke, M. E., J. G. Babb, and M. K. Ritke. 1991. Breeding-site specificity in the gray treefrog (HYLA CHRYSOSCELIS). J. Herpetol. 25:123-125.
  24. Vogt, R. C. 1981c. Natural history of amphibians and reptiles of Wisconsin. Milwaukee Public Museum. 205 pp.
  25. Wiley, J. E. 1983. Chromosome polymorphism in HYLA CHRYSOSCELIS. Copeia 1983:273-275.