Antrozous pallidus

(Le Conte, 1856)

Pallid Bat

G4Apparently Secure Found in 38 roadless areas NatureServe Explorer →
G4Apparently SecureGlobal Rank
Least concernIUCN
MediumThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.106431
Element CodeAMACC10010
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassMammalia
OrderChiroptera
FamilyVespertilionidae
GenusAntrozous
Other Common Names
Chauve-souris blonde (FR) Un Murciélago (ES)
Concept Reference
Wilson, D. E., and D. M. Reeder (editors). 1993. Mammal species of the world: a taxonomic and geographic reference. Second edition. Smithsonian Institution Press, Washington, DC. xviii + 1206 pp. Available online at: http://www.nmnh.si.edu/msw/.
Taxonomic Comments
Although Antrozous koopmani of Cuba is treated as a separate species by ASM (2025) and Simmons and Cirranello (2025), both sources state that additional research is needed. It is possibly extinct and only known from a few modern and fossil representatives.
Conservation Status
Rank MethodLegacy Rank calculation - Excel v3.1x
Review Date2016-04-04
Change Date2015-03-24
Edition Date2015-03-24
Edition AuthorsHammerson, G.
Threat ImpactMedium
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Rank Reasons
Large range in western North America; in some areas, habitat has been reduced and degraded by agricultural expansion, cheatgrass invasion, fire, urban development, excessive livestock grazing, and pesticide use, but the species remains widespread, common, and likely relatively stable in many areas of the arid southwestern United States; however, regional population trends are poorly known.
Range Extent Comments
Range includes western North America from south-central British Columbia (Okanagan Valley; small resident population; Willis and Bast 2000; Rambaldini and Brigham 2008, 2011) south through the western United States to southern Baja California, central Mexico, southern Kansas, and southern Texas; also Cuba (Martin and Schmidly 1982, Hermanson and O'Shea 1983, Reid 2006). Elevational range in Texas is 600-2,000 meters (Ammerman et al. 2012).

The following subspecies distributions are from Martin and Schmidly (1982). Subspecies pacificus: Pacific Coast Ranges of western Oregon and California south to Los Angeles and San Bernardino counties. Subspecies pallidus: east of the range of pacificus from southern British Columbia and east of the Cascade Range throughout much of the Columbia Plateau and Great Basin, throughout the southwestern U.S.. west of central Texas, and south to western and south-central Mexico north of the Transverse Volcanic Cordillera. Subspecies bunkeri: Barber County, Kansas, south to the western end of the Wichita Mountains in Greer County, Oklahoma. Subspecies minor: s. Baja California north through the Colorado Desert of southeastern California and southwestern Arizona, thence northward into southern Nevada. Subspecies packardi: western slopes of the Sierra Madre Occidental in southwestern Zacatecas, Jalisco, northeastern Nayarit, and southern Sonora. Subspecies koopmani: several scattered localities in Cuba.
Occurrences Comments
The number of distinct occurrences has not been determined using standardized criteria, but this species is represented by a large number of collection/observation sites and locations (as defined by IUCN).
Threat Impact Comments
On a range-wide basis, no major threats are known. Locally, some maternity colonies and hibernacula are susceptible to disturbance, and they may be negatively affected or destroyed as a result of vandalism, mine closures or reactivation, or other activities. Tree-roosting populations (Baker et al. 2008) may be detrimentally affected by timber harvest and other forestry practices. Roosts in buildings may be lost as a result of demolition, bat exclusion, or other alterations. Some populations undoubtedly have been negatively affected by loss or extensive modification of primary foraging habitat caused by agricultural expansion (including orchards and vineyards), cheatgrass invasion, fire, urban development, excessive livestock grazing, and pesticide use (Willis and Bast 2000), but the degree of impact of these threats on the affected populations is not well known.

Apparent declines in lowland south-coastal California likely are associated with loss of habitat primarily due to urban expansion (Miner and Stokes 2005).

The species undoubtedly has lost considerable habitat in Washington because of agricultural expansion (Hayes and Wiles 2013).

As of mid-2012, this species was not known to be affected by white-nose syndrome.
Ecology & Habitat

Description

A large, pale bat with large ears (not joined at base), large eyes, a simple muzzle, and yellowish drab dorsal pelage (palest in deserts, darkest along coast); total length 92-135 mm; tail length 35-53 mm; hind foot length 11-16 mm; ear length 21-37 mm; forearm length 45-60 mm; skull length 18.6-24 mm; mass 13.6-24.1 g in males, 13.9-28.0 g in females (Hermanson and O'Shea 1983).

Diagnostic Characteristics

Differs from most other vespertilionids in having much larger ears, larger eyes, and paler pelage. Differs from big-eared bats (CORYNORHINUS) in having ears that are not joined at the base. Differs from Macrotus bats in lacking a prominent noseleaf.

Habitat

Habitats include mountainous areas, intermontane basins, and lowland desert scrub (Ammerman et al. 2012); arid deserts and grasslands (Adams 2003), often near rocky outcrops and water; in some areas, this species also inhabits open coniferous forest and woodland (Baker et al. 2008). In British Columbia, pallid bats foraged over native habitats and also over vineyards (Rambaldini and Brigham 2011).

Day roosts include crevices of rock outcrops, caves, mine tunnels, buildings, bridges, and hollows of live and dead trees. Night roosts often or typically are in caves in Oklahoma (Caire et al. 1989). In Oregon, night roosts were in buildings, under rock overhangs, and under bridges; bats generally were faithful to particular night roosts both within and between years (Lewis 1994).

Hibernation occurs in caves and mines, though not very many hibernation records are available.

Young are born in maternity colonies usually in rock crevices or buildings.

Ecology

This is a gregarious species. Usually it forms clusters in diurnal roosts. It may gather in night roosts that are frequently near, but separate from, day roosts (Lewis 1994). Tadarida brasiliensis and Myotis yumanensis may roost among pallid bats.

This species is a legitimate pollinator of certain agaves and cacti (Herrera et al. 1993, Frick et al. 2014).

Reproduction

Copulation usually occurs in October-December. Delayed fertilization in spring. In the U.S., young are born in late May-early June in California, mostly late June in Kansas, probably early May to mid-June in Texas (Schmidly 1991). Litter size usually is 2, sometimes 1. Young fly at 6 weeks, weaned in 6-8 weeks (Hermanson and O'Shea 1983). In Oregon, reproductive success was reduced in a year with low spring temperatures (Lewis 1993). Maternity colonies usually are small, but may include up to 200+ adults, may include adult males.
Terrestrial Habitats
Woodland - ConiferWoodland - MixedShrubland/chaparralGrassland/herbaceousDesertCliffSuburban/orchard
Palustrine Habitats
Riparian
Other Nations (2)
CanadaN2
ProvinceRankNative
British ColumbiaS2Yes
United StatesN5
ProvinceRankNative
IdahoS3Yes
UtahS4Yes
NevadaS3Yes
ColoradoS4Yes
WyomingS2BYes
CaliforniaS3Yes
WashingtonS3Yes
ArizonaS4Yes
New MexicoS4Yes
OklahomaS3Yes
MontanaS3Yes
Navajo NationS5Yes
OregonS2Yes
KansasS1Yes
TexasS5Yes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentNegligible (<1%)High (continuing)
2 - Agriculture & aquacultureSmall (1-10%)Moderate - slightHigh (continuing)
2.3 - Livestock farming & ranchingSmall (1-10%)Moderate - slightHigh (continuing)
3 - Energy production & miningSmall (1-10%)Extreme or 71-100% pop. declineHigh (continuing)
3.2 - Mining & quarryingHigh (continuing)
3.3 - Renewable energyHigh (continuing)
4 - Transportation & service corridorsNegligible (<1%)High (continuing)
5 - Biological resource useNegligible (<1%)High (continuing)
6 - Human intrusions & disturbanceSmall (1-10%)Moderate - slightHigh (continuing)
6.1 - Recreational activitiesSmall (1-10%)Extreme - moderateHigh (continuing)
6.3 - Work & other activitiesSmall (1-10%)Extreme - moderateHigh (continuing)
7 - Natural system modificationsNegligible (<1%)High (continuing)
7.2 - Dams & water management/useHigh (continuing)
8 - Invasive & other problematic species, genes & diseasesRestricted (11-30%)Moderate - slightHigh (continuing)
9 - PollutionUnknownUnknownHigh (continuing)
10 - Geological eventsNegligible (<1%)
11 - Climate change & severe weatherPervasive (71-100%)UnknownHigh (continuing)

Roadless Areas (38)
Arizona (10)
AreaForestAcres
Catalina St. Pk. Roadless AreaCoronado National Forest951
ChiricahuaCoronado National Forest76,876
GaliuroCoronado National Forest28,333
Jacks CanyonCoconino National Forest2,858
MazatzalTonto National Forest16,942
Middle Romero WSRCoronado National Forest60
Oracle RoadlessCoronado National Forest22,365
Sierra Ancha Wilderness ContiguousTonto National Forest7,787
TumacacoriCoronado National Forest44,594
WhetstoneCoronado National Forest20,728
California (13)
AreaForestAcres
AntimonyLos Padres National Forest40,911
Bell QuinbyShasta-Trinity National Forest11,556
Cow CreekShasta-Trinity National Forest22,627
Coyote NorthInyo National Forest11,932
Dennison PeakSequoia National Forest6,293
Devil GulchSierra National Forest30,490
Fox MountainLos Padres National Forest52,072
Grouse LakesTahoe National Forest19,085
La BreaLos Padres National Forest14,031
Little French CShasta-Trinity National Forest11,529
San JoaquinSierra National Forest22,474
Santa CruzLos Padres National Forest21,182
South ForkShasta-Trinity National Forest16,786
Nevada (4)
AreaForestAcres
Georges CanyonHumboldt-Toiyabe National Forest108,551
Pine Grove SouthHumboldt-Toiyabe National Forest88,945
SaulsburyHumboldt-Toiyabe National Forest30,957
Warm SpringsHumboldt-Toiyabe National Forest29,540
New Mexico (8)
AreaForestAcres
Candian RiverCibola National Forest7,149
Capitan MountainsLincoln National Forest14,069
Contiguous To Black & Aldo Leopold WildernessGila National Forest111,883
Devils CreekGila National Forest89,916
Dry CreekGila National Forest26,719
Gila BoxGila National Forest23,759
Last Chance CanyonLincoln National Forest8,934
Meadow CreekGila National Forest34,167
Utah (3)
AreaForestAcres
Clarkston Mtn.Caribou National Forest7,099
CottonwoodDixie National Forest6,754
Pine Valley MountainsDixie National Forest57,673
References (52)
  1. Adams, R. A. 2003. Bats of the Rocky Mountain West: natural history, ecology, and conservation. University Press of Colorado, Boulder, Colorado. xiii + 289 pp.
  2. American Society of Mammalogists (ASM). 2025. Mammal Diversity Database (Version 1.13) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.10595931. Online. Available: https://www.mammaldiversity.org/
  3. Ammerman, L. K., C. L. Hice, and D. J. Schmidly. 2012. Bats of Texas. Texas A &amp; M University Press, College Station, Texas. xvi + 305 pp.
  4. Arita, H. T. 1993. Conservation biology of the cave bats in Mexico. Journal of Mammalogy 74:693-702.
  5. Baker, M. D., M. J. Lacki, G. A. Falxa, P. L. Droppelman, R. A. Slack, and S. A. Slankard. 2008. Habitat use of pallid bats in coniferous forests of northern California. Northwest Science 82:269-275.
  6. Balcombe, J. P. 1988c. Status report on the Pallid Bat <i>Antrozous pallidus</i> in Canada. Committee on the Status of Endangered Wildlife in Canada (COSEWIC). 13 pp.
  7. Banfield, A. W. F. 1974. The mammals of Canada. University of Toronto Press, Toronto, Canada. 438 pp.
  8. Barbour, R. W., and W. H. Davis. 1969. Bats of America. The University of Kentucky Press, Lexington, Kentucky. 286 pp.
  9. Bradley, R.D., L.K. Ammerman, R.J. Baker, L.C. Bradley, J.A. Cook. R.C. Dowler, C. Jones, D.J. Schmidly, F.B. Stangl Jr., R.A. Van den Bussche and B. Würsig. 2014. Revised checklist of North American mammals north of Mexico, 2014. Museum of Texas Tech University Occasional Papers 327:1-28. Available at: http://www.nsrl.ttu.edu/publications/opapers/ops/OP327.pdf
  10. Brown, P. E., R. D. Berry, K. L. Miner, and H. Johnson. 1997. Roosting behavior of pallid bats <i>Antrozous pallidus</i> in the California desert as determined by radio-telemetry. Bat Research News 38:100.
  11. Caire, W., J. D. Tyler, B. P. Glass, and M. A. Mares. 1989. Mammals of Oklahoma. University of Oklahoma Press, Norman. Oklahoma. 567 pp.
  12. Clawson, R. L., R. K. LaVal, M. L. LaVal, and W. Claire. 1980. Clustering behavior of hibernating <i>Myotis sodalis</i> in Missouri, USA. Journal of Mammalogy 61:245-53.
  13. Davis, R. 1966. Homing performance and homing ability in bats. Ecological Monographs 36:201-237.
  14. Engstrom, M. D., and D. E. Wilson. 1981. Systematics of ANTROZOUS DUBIAQUERCUS (Chiroptera: Vespertilionidae), with comments on the status of BAUERUS Van Gelder. Annals of Carnegie Museum 50:371-383.
  15. Fellers, G. M., and E. D. Pierson. 2002. Habitat use and foraging behavior of Townsend's Big-eared Bat (<i>Corynorhinus townsendii</i>) in coastal California. Journal of Mammalogy 83:167-177.
  16. Frick, W. F., J. R. Shipley, J. F. Kelly, P. A. Heady, III, and K. M. Kay. 2014. Seasonal reliance on nectar by an insectivorous bat revealed by stable istopes. Oecologia 174:55-65.
  17. Garner, J. D., and J. E. Gardner. 1992. Determination of summer distribution and habitat utilization of the Indiana bat (<i>Myotis sodalis</i>) in Illinois. Illinois Department of Conservation. Final Report, Project E-3. Springfield, IL, 23 pp.
  18. Griffin, D. R. 1940b. Migrations of New England bats. Bulletin of the Museum of Comparative Zoology 86:217-246.
  19. Hall, E. R. 1981a. The Mammals of North America, second edition. Vols. I &amp; II. John Wiley &amp; Sons, New York, New York. 1181 pp.
  20. Hayes, G., and G. J. Wiles. 2013. Washington bat conservation plan. Washington Department of Fish and Wildlife, Olympia, Washington. viii + 138 pp.<br>
  21. Hermanson, J. W., and T. J. O'Shea 1983. <i>Antrozous pallidus</i>. Mammalian Species 213:1-8.
  22. Herrera, L. G., T. H. Fleming, and J. S. Findley. 1993. Geographic variation in carbon composition of the pallid bats, <i>Antrozous pallidus</i>, and its dietary implications. Journal of Mammalogy 74:601-606.
  23. Jones, J. K., Jr., R. S. Hoffman, D. W. Rice, C. Jones, R. J. Baker, and M. D. Engstrom. 1992a. Revised checklist of North American mammals north of Mexico, 1991. Occasional Papers, The Museum, Texas Tech University, 146:1-23.
  24. Kiser, J. D. and C. L. Elliott. 1996. Foraging habitat, food habits, and roost tree characteristics of the Indiana bat (<i>Myotis sodalis</i>) during autumn in Johnson County, Kentucky. Final report, Kentucky Department of Fish and Wildlife Resources, Frankfort, Kentucky. 65pp.
  25. Lewis, S. E. 1993. Effect of climatic variation on reproduction by pallid bats (ANTROZOUS PALLIDUS). Can. J. Zool. 71:1429-1433.
  26. Lewis, S. E. 1994b. Night roosting ecology of pallid bats (<i>Antrozous pallidus</i>) in Oregon. Am. Midl. Nat. 132:219-226.
  27. Linzey, D. W. 1998. The mammals of Virginia. McDonald and Woodward, Blacksburg, VA. 459 pp.
  28. Mammalian Species, nos. 1-604. Published by the American Society of Mammalogists.
  29. Manning, R. W., et al. 1988. Subspecific status of the pallid bat, ANTROZOUS PALLIDUS, in the Texas Panhandle and adjacent areas. Occas. Pap. Mus. Texas Tech Univ. 118:1-5.
  30. Martin, C. O., and D. J. Schmidly. 1982. Taxonomic review of the pallid bat, Antrozous pallidus (Le Conte). Spec. Publ. Mus., Texas Tech. Univ. 18:1-48.
  31. Menzel, M. A., R. Odom, S. Owen, W. M. Ford, B. R. Chapman, K. V. Miller, J. Edwards, and P. Wood. 1999b. Investigation of foraging habitat use by bats with a focus on Northern Long-eared Myotis (<i>Myotis septentrionalis</i>): a comparison of methods. IN M. K. Clark, editor. Abstracts from the 1999 Colloquium on the conservation of mammals in the Southeastern United States. Available at: http://www.batworkinggroups.org/sbdnnews.htm. Accessed 2001-06-12.
  32. Miner, K. L., and D. C. Stokes. 2005. Bats in the South Coast Ecoregion: status, conservation issues, and research needs. Pages 211-227 in USDA Forest Service General Technical Report PSW-GTR-195.
  33. Morgan, C. N., L. K. Ammerman, K. D. Demere, J. B. Doty, Y. J. Nakazawa, and M. R. Mauldin. 2019. Field identification key and guide for bats of the United States of America. Occasional Papers, Museum of Texas Tech University, Number 360. Texas Tech University Natural Science Research Laboratory. 29 pp.
  34. Oliver, G. V. 2000. The bats of Utah: a literature review. Publication No. 00-14, Utah Division of Wildlife Resources, Salt Lake City.
  35. Orihuela, J., L. Pérez Orozco, J. L. Álvarez Licourt, R. A. Viera Muñoz, and C. Santana Barani. 2020a. Late Holocene land vertebrate fauna from Cueva de los Nesofontes, Western Cuba: stratigraphy, chronology, diversity, and paleoecology. Palaeontologia Electronica 23(3):a57.
  36. Orihuela, J., L. W. Viñola, and R. A. Viera. 2020b. New bat locality records from Cuba with emphasis on the province of Matanzas. Novitates Caribaea 15:96-116.
  37. Ports, M. A., and P. V. Bradley. 1996. Habitat affinities of bats from northeastern Nevada. Great Basin Naturalist 56:48-53.
  38. Rambaldini, D., A., and R. M. Brigham. 2008. Torpor use by free-ranging pallid bats (<i>Antrozous pallidus</i>) at the northern extent of their range. Journal of Mammalogy 89(4): 933-941
  39. Rambaldini, D., A., and R. M. Brigham. 2011. Pallid bat (<i>Antrozous pallidus</i>) foraging over native and vineyard habitats in British Columbia, Canada. Canadian Journal of Zoology 89(9):816-822.
  40. Ransome, R. 1990. The natural history of hibernating bats. Christopher Helm, London. xxi + 235 pp.
  41. Reid, F. A. 2006. A field guide to mammals of North America north of Mexico. Houghton Mifflin Company, Boston.
  42. Rodríguez-Durán, A. and T.H. Kunz. 2001. Biogeography of West Indian bats: An ecological perspective. Pp. 355-368 In C. A. Woods and F. E. Sergile (eds.), Biogeography of the West Indies. CRC Press, Boca Raton, FL, USA.
  43. Schmidly, D. J. 1977. The mammals of Trans-Pecos Texas including Big Bend National Park and Guadalupe Mountains National Park. Texas A & M University Press, College Station.
  44. Schmidly, D. J. 1991. The bats of Texas. Texas A & M University Press, College Station, Texas. 188 pp.
  45. Simmons, N. B. and A. L. Cirranello. 2025. Bat Species of the World: A taxonomic and geographic database. Version 1.7. Online. Available: https://batnames.org/
  46. Tuttle, M. D. 1976a. Population ecology of the gray bat (<i>Myotis grisescens</i>): philopatry, timing, and patterns of movement, weight loss during migration, and seasonal adaptive strategies. University of Kansas Museum of Natural History Occasional Papers (54):1-38.
  47. U.S. Fish and Wildlife Service (USFWS). 1999. Agency draft Indiana Bat (<i>Myotis sodalis</i>) revised recovery plan. U.S. Fish and Wildlife Service, Fort Snelling, Minnesota. 53 pp.
  48. van Zyll de Jong, C.G. 1985. Handbook of Canadian Mammals. Vol. II, Bats. National Museum of Natural Sciences, National Museums of Canada, Ottawa, Canada. 212 pp.
  49. Willis, C.K.R., and M. L. Bast. 2000. COSEWIC assessment and update status report on the pallid bat <i>Antrozous pallidus</i> in Canada. Committee on the Status of Endangered Wildlife in Canada, Ottawa, Ontario.<br />
  50. Wilson, D. E., and D. M. Reeder (editors). 1993. Mammal species of the world: a taxonomic and geographic reference. Second edition. Smithsonian Institution Press, Washington, DC. xviii + 1206 pp. Available online at: http://www.nmnh.si.edu/msw/.
  51. Wilson, D. E., and D. M. Reeder (editors). 2005. Mammal species of the world: a taxonomic and geographic reference. Third edition. The Johns Hopkins University Press, Baltimore. Two volumes. 2,142 pp. [As modified by ASM the Mammal Diversity Database (MDD) at https://www.mammaldiversity.org/index.html]
  52. Wilson, D. E., and S. Ruff. 1999. The Smithsonian book of North American mammals. Smithsonian Institution Press, Washington, D.C. 750 pp.