Range Extent CommentsUpper and middle Mobile drainage, Alabama, Georgia, and southeastern Tennessee (absent from Tombigbee River system); uplands and outliers of southern Ohio River basin and Ozarks, west to eastern Oklahoma, south to northern Alabama and northern Georgia, east to western Virginia, western North Carolina; northern Ohio River basin and north into middle part of upper Mississippi River basin to southeastern Minnesota, southern Wisconsin (Fago and Hauber 1993), Pennsylvania, and West Virginia; southern Great Lakes basin, north to western New York, southern Ontario, Michigan, and southern Wisconsin. Avoids lowlands of central Mississippi River basin. See Parker (1989) for information on status in Canada.