Habitat
Alpine meadows in south; mountain valleys in north. Wet meadows, cropland, especially fields & pastures of grass and legumes along fence rows; grassy areas by streams, lakes. Occupies shallow burrows and surface runways.
Ecology
Peak popualtion density was 375-560/ha in Utah (Negus et al. 1986). Populations may fluctuate dramatically. Populations peaked at 3-4-year intervals in northwestern Wyoming (Pinter 1986). Predators include hawks, owls, foxes, badgers, coyotes, etc.
Reproduction
Breeds April-October (births April-August in Utah, Negus et al. 1986; May-August in northwestern Wyoming, Negus et al. 1992). Usually 2-3 litters/year (4 cohorts/year in Utah, 3/year in northwestern Wyoming; early cohorts breed in same season). Average litter size is about 6; litter size peaked at 3-4-year intervals in northwestern Wyoming (Pinter 1986). Females of early cohorts begin breeding at 4-5 weeks in favorable years. Drought greatly reduced/delayed growth and attainment of sexual maturity in northwestern Wyoming (Negus et al. 1992).