Oreohelix strigosa

(Gould, 1846)

Rocky Mountainsnail

G5Secure Found in 37 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Identity
Unique IDELEMENT_GLOBAL.2.109795
Element CodeIMGASB5320
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryInvertebrate Animal
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumMollusca
ClassGastropoda
OrderStylommatophora
FamilyOreohelicidae
GenusOreohelix
Synonyms
Patula strigosa(A. Gould, 1846)
Concept Reference
Turgeon, D. D., J. F. Quinn, Jr., A. E. Bogan, E. V. Coan, F. G. Hochberg, W. G. Lyons, P. M. Mikkelsen, R. J. Neves, C. F. E. Roper, G. Rosenberg, B. Roth, A. Scheltema, F. G. Thompson, M. Vecchione, and J. D. Williams. 1998. Common and scientific names of aquatic invertebrates from the United States and Canada: Mollusks. 2nd Edition. American Fisheries Society Special Publication 26, Bethesda, Maryland. 526 pp.
Taxonomic Comments
Considerable confusion exists in the literature, especially in older works, concerning Oreohelix strigosa and Oreohelix subrudis, which were frequently confused, O. subrudis having often been included in O. strigosa, at least until the mid-1930s. Older literature is further complicated by use of the name "Oreohelix cooperi", formerly applied variously to both O. strigosa and O. subrudis; the name cooperi has also been used for various races of each of both species, but, as used now, this name has been restricted to one race of O. strigosa.

Although Brandauer (1988), based on both external (shell) and internal (genitalia and radula) morphology, concluded that Oreohelix subrudis should be considered only a form (or "morph") of Oreohelix strigosa (i.e., that the two are synonyms), Rees (1988) found that "[c]onsistent and distinct differences in enzyme banding patterns between these two species were revealed by electrophoresis" and "allozyme variation can be assessed to reliably distinguish between O. strigosa and O. subrudis."

Pilsbry (1939) discussed 10 subspecies and numerous "forms" and "varieties" of Oreohelix strigosa, but his subspecies concept was very different from the one that is in current taxonomic use, many of Pilsbry's subspecies now being regarded as mere "morphs". Richardson (1984) listed 25 subspecies (including one fossil race) of Oreohelix strigosa. It is questionable whether all of these subspecies merit recognition, for O. strigosa is one of the most plastic and polymorphic of all gastropods, exhibiting surprisingly great variation even within local populations.

Weaver et al. (2008) concluded that specimens identified as Oreohelix peripherica wasatchensis are not monophyletic based on molecular and radular data; and instead show two distinct clades within "O. p. wasatchensis", one that nests within Oreohelix strigosa, and one that is a sister group to Oreohelix peripherica peripherica. Their evidence indicates Oreohelix strigosa may not be a monophyletic group and add that taxonomic revision of the species is necessary.
Conservation Status
Review Date2002-10-08
Change Date2002-10-08
Edition Date1998-08-31
Edition AuthorsG. V. Oliver
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences81 to >300
Rank Reasons
Widespread, abundant, and, within its range, apparently one of the most secure of all mollusk species, including exotic introductions.
Range Extent Comments
Occurs generally throughout the mountainous areas of interior western North America. Known from southeastern British Columbia, extreme southeastern Alberta, eastern Washington, extreme northeastern Oregon, northern and extreme southeastern Idaho, western Montana, extreme western South Dakota (i.e., the Black Hills), extreme western Wyoming, extreme eastern Nevada, nearly all of Utah, Colorado except the eastern third, northern and eastern Arizona, and northern and central New Mexico (Pilsbry 1939; see also Bequaert and Miller 1973). Known also as a Pleistocene fossil in Iowa and Illinois (Pilsbry 1939, Morrison 1943, Frest and Rhodes 1981). Pilsbry (1939) commented on the discontinuities in the northern and northwestern parts of its range (i.e., Washington and Oregon). Disjunct populations are also apparent at the northeastern and southern limits of distribution, some notable examples of such disjunct--probably Pleistocene relictual--populations being those in the Black Hills (South Dakota), the White Mountains (Arizona), and the Sierra Blanca (New Mexico).
Occurrences Comments
Certainly well over 100 and possibly 1,000 or more extant occurrences. Pilsbry's dot map (1939, Figure 296) shows 106 specimen localities (for this species), and very many more are known. For example, in Utah alone only a small fraction of the extant occurrences are represented in Pilsbry's (1939) map. Distribution in Alberta is spotty: Cypress Hills with the next nearest location being Waterton Lake, then north of Pincher Creek (Lepitzki, 2001). Most recently, it was discovered in the Ktunaxa Traditional Territory in southeastern British Columbia (which extends from near Canada - U.S. border north to about 50 km north of Cranbrook) (Ovaska and Sopuck, 2009).
Threat Impact Comments
Generally protected from most anthropogenic threats by the unsuitablity of its habitat for development and other economic use; however, timber harvest within its habitat is very likely a threat. Fire represents a natural threat.
Ecology & Habitat

Habitat

Composition of the plant community appears to be of little importance, dominant plant species where it occurs ranging from sagebrush to a wide variety of deciduous shrubs and trees and a similarly wide variety of coniferous shrubs and trees; reported habitats, described in general terms, vary greatly and range from "chaparral" to forest and barren rock slides. Substrate, however, is of great importance, the presence of exposed limestone being almost critical for occurrence; exceptions, however, are well known, there being documented occurrences on sandstone, and occurrences on other substrates probably exist. Slope, too, has been considered to be of importance. Jones (1940) considered the presence of "moisture, lime[stone], and leaf mould" to be of critical importance, colonies being "found chiefly in or near limestone debris at the base of shady cliffs."

Elevational range great: Pilsbry (1939) mentioned localities as low as 800 to 900 feet in Idaho and as high as 10,000 feet in Utah and possibly Montana, and Hoff (1962) reported it at 11,800 feet in New Mexico.
Other Nations (2)
United StatesN5
ProvinceRankNative
New MexicoSNRYes
NevadaSNRYes
WyomingSUYes
MontanaS5Yes
ArizonaS4Yes
UtahS5Yes
IllinoisSNRYes
IowaSXYes
IdahoSNRYes
OregonSNRYes
South DakotaSNRYes
WashingtonS4Yes
Navajo NationSNRYes
CanadaN4
ProvinceRankNative
British ColumbiaS4Yes
Roadless Areas (37)
Idaho (1)
AreaForestAcres
PalisadesCaribou-Targhee National Forest122,002
Montana (1)
AreaForestAcres
Big Snowy Mountains WsaLewis and Clark National Forest88,003
Nevada (9)
AreaForestAcres
Moriah - Rye Grass BHumboldt-Toiyabe National Forest392
Ruby - Cold CkHumboldt-Toiyabe National Forest2,339
Ruby - Lamoille CynHumboldt-Toiyabe National Forest32,771
Ruby - Secret Pk.Humboldt-Toiyabe National Forest6,749
Ruby - ThompsonHumboldt-Toiyabe National Forest5,289
Snake - Big WashHumboldt-Toiyabe National Forest4,146
Snake - ChokecherryHumboldt-Toiyabe National Forest30,845
Snake - MurphyHumboldt-Toiyabe National Forest27,064
South SchellHumboldt-Toiyabe National Forest125,614
New Mexico (6)
AreaForestAcres
Black CanyonSanta Fe National Forest1,922
Columbine - Hondo Wilderness Study AreaCarson National Forest43,739
Holy GhostSanta Fe National Forest2,352
Jacks CreekSanta Fe National Forest740
Pecos WS RiverSanta Fe National Forest5,396
Tesuque CreekSanta Fe National Forest810
Utah (11)
AreaForestAcres
0401005Ashley National Forest38,930
418017Uinta National Forest19,631
Clarkston Mtn.Caribou National Forest7,099
East MountainManti-Lasal National Forest30,705
Horse Mountain - Mans PeakManti-Lasal National Forest22,159
Mt. AireWasatch-Cache National Forest9,681
Mt. Logan NorthWasatch-Cache National Forest18,930
Mt. NaomiWasatch-Cache National Forest41,922
Right Hand Fork LoganWasatch-Cache National Forest15,023
South FrancisWasatch-Cache National Forest3,374
Stansbury MountainsWasatch-Cache National Forest39,696
Wyoming (9)
AreaForestAcres
Cloud Peak ContiguousBighorn National Forest113,757
Gannett Hills - Spring CreekBridger-Teton National Forest45,462
Grayback RidgeBridger-Teton National Forest295,113
Lake Alice - Commissary RidgeBridger-Teton National Forest166,707
Leigh CreekBighorn National Forest19,180
PalisadesTarghee National Forest1,121
Salt River RangeBridger-Teton National Forest235,661
South Wyoming RangeBridger-Teton National Forest85,776
Wilderness Study AreaTarghee National Forest51,961
References (21)
  1. Beetle, D.E. 1957. The Mollusca of Teton County, Wyoming. Nautilus 71:12-22.
  2. Bequaert, J.C. and W.B. Miller. 1973. The mollusks of the arid southwest with an Arizona checklist. University of Arizona Press: Tucson, Arizona. 271 pp.
  3. Brandauer, N.E. 1988. Family Oreohelicidae (Gastropoda: Pulmonata) in Colorado. Natural History Inventory of Colorado 9:1-32.
  4. Chamberlin, R.V. and D.T. Jones. 1929. A descriptive catalog of the Mollusca of Utah. Bulletin of the University of Utah, [Biological Series 1(1)] 19(4): 1-203.
  5. Frest, T.J. and R.S. Rhodes, II. 1981. <i>Oreohelix strigosa cooperi </i>(Binney) in the Midwest Pleistocene. The Nautilus, 95: 47-55.
  6. Gregg, W.O. 1940. Mollusca of Zion National Park, Utah. Nautilus 54:30-32.
  7. Gregg, W.O. 1941. Mollusca of Cedar Breaks National Monument, Utah. Nautilus 54:116-117.
  8. Henderson, J. 1953. Mollusca of the Yellowstone Park, Teton Park and Jackson Hole region. Nautilus 47:1-3.
  9. Hoff, C.C. 1962. Some terrestrial Gastropoda from New Mexico. Southwestern Naturalis 7:51-63.
  10. Jones, D.T. 1940. A study of the Great basin land snail OREOHELIX STRIGOSA DEPRESSA (Cockerell). Bulletin of the University of Utah [Biological Series 6(1)] 31(4):1-43.
  11. Lepitzki, D.A.W. 2001. Gastropods: 2000 preliminary status ranks for Alberta. Unpublished report prepared for Alberta Sustainable Resource Development, Fish and Wildlife Division, Edmonton, Alberta. 126 pp.
  12. Morrison, J.P.E. 1943. OREOHELIX east of the Mississippi. Nautilus 56:104.
  13. Ovaska, K. and L. Sopuck. 2009. Surveys for terrestrial gastropods at risk within Ktunaxa Traditional Territory. Report prepared for British Columbia Ministry of Environment, Victoria, British Columbia. 27 pp.
  14. Pilsbry, H. A. 1939a-1948. Land Mollusca of North America (North of Mexico.) George W. Carpenter Fund for the Encouragement of Original Scientific Research, Philadelphia, Pennsylvania. 2107 pp.
  15. Rees, B.B. 1988. Electrophoretic and morphological characteristics of two species of OREOHELIX, the mountain snail. Malacological Review 21:129-132.
  16. Richardson, L. 1984. Oreohelicidae: catalog of species. Tryonia 10:I+1-30.
  17. Spamer, E.E., and A.E. Bogan. 1993. Mollusca of the Grand Canyon and vicinity, Arizona; new and revised data on diversity and distributions, with notes on Pleistocene-Holocene mollusks of the Grand Canyon. Proceedings of the Academy of Natural Sciences of Philadelphia 144:21-68.
  18. Turgeon, D. D., J. F. Quinn, Jr., A. E. Bogan, E. V. Coan, F. G. Hochberg, W. G. Lyons, P. M. Mikkelsen, R. J. Neves, C. F. E. Roper, G. Rosenberg, B. Roth, A. Scheltema, F. G. Thompson, M. Vecchione, and J. D. Williams. 1998. Common and scientific names of aquatic invertebrates from the United States and Canada: Mollusks. 2nd Edition. American Fisheries Society Special Publication 26, Bethesda, Maryland. 526 pp.
  19. Vanatta, E.G. 1921. Shells of Zion National Park, Utah. Nautilus 34:140-141.
  20. Weaver, K.F., M. Perez-Losada, R.P. Guralnick, A. Nelson, S. Blatt, and K.A. Crandall. 2008. Assessing the conservation status of the land snail <i>Oreohelix peripherica wasatchensis</i> (Family Oreohelicidae). Conservation Genetics, 9: 907-916.
  21. Woodbury, A.M. 1929. The snails of Zion National Park. Nautilus 43:54-61.