Trillium albidum

J.D. Freeman

White Toadshade

G4Apparently Secure (G4G5) Found in 41 roadless areas NatureServe Explorer →
G4Apparently SecureGlobal Rank
Least concernIUCN
Medium - lowThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.1128612
Element CodePMLIL201E0
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVascular Plant
IUCNLeast concern
Endemicendemic to a single nation
KingdomPlantae
PhylumAnthophyta
ClassMonocotyledoneae
OrderLiliales
FamilyMelanthiaceae
GenusTrillium
Concept Reference
Meyers, S.C., T. Jaster, K. E. Mitchell, and L.K. Hardison, editors. 2015. Flora of Oregon. Volume 1: Pteridophytes, Gymnosperms, and Monocots. Botanical Research Institute of Texas, Fort Worth, TX. 608 pp.
Taxonomic Comments
This species is made up of one subspecies parviflorum that occurs in Washington south to northern Oregon where it intergrades with the typical subspecies (Flora of the Pacific Northwest 2018).
Conservation Status
Rank Method Rank calculation - Biotics v2
Review Date2019-10-23
Change Date2019-09-13
Edition Date2019-09-20
Edition AuthorsOliver, L. and K. Wayman
Threat ImpactMedium - low
Range Extent200,000-2,500,000 square km (about 80,000-1,000,000 square miles)
Rank Reasons
Trillium albidum is comprised of the typical subspecies and subspecies parviflorum. The range of the species is from the western Cascades south through coastal California. Subspecies parviflorum is threatened by loss of habitat and invasive species. The typical subspecies is primarily threatened by wildfires.
Range Extent Comments
Trillium albidum occurs from the western Cascades in Pierce County Washington south to California (Hitchcock and Cronquist 2018).
Threat Impact Comments
The subspecies parviflorum, in the northern portion of the range, is threatened by habitat conversion and fragmentation, recreation use, development, elk herbivory, and by non-native plants. Wildfire is a particular threat to subspecies albidum.
Ecology & Habitat

Reproduction

Trillium seeds have an elaiosome, an oily, lipid-rich attachment that is highly attractive to ants. The ants carry the seeds to their nest, eat the attachment, and leave the seeds in tunnels in their nests (FNA 2002a, Leege et al. 2010). The seeds later germinate en masse (Case and Case 1997). Yellow jackets (Vespula spp.) and other wasps are similarly attracted to the elaiosome. Yellow jackets are documented seed dispersers for three species (T. catesbaei, T. cuneatum, T. undulatum) (Zettler et al. 2001). Ants carry the seeds an average of about 1m whereas yellow jackets disperse seeds an average of 1.4m (Chafin 2010, Zettler et al. 2001). Long distance dispersers include mammals, such as white-tailed deer (Odocoileus virginianus) and woodchucks (Marmota monax) (Chafin 2010, Vellend et al. 2006). Similarly, it is suspected that elk (Cervus canadensis) and mule deer (Odocoileus hemionus) of the western U.S.A. are capable of dispersing seeds over long distances (Bartuszevige and Endress 2008).

All Trillium have rhizomes but the frequency of asexually reproduction varies (Chauhan et al. 2019, FNA 2002a, Ohara 1989).

Trillium seeds exhibit a somewhat unique kind of dormancy called deep simple double morphophysiological dormancy, meaning they require two winters and one summer to complete dormancy break. After dispersal, roots (radicles) emerge in the first spring and leaves (epicotyls) begin growing in the second spring. The result is that Trillium seeds are generally about 1.5-2 years before they are non-dormant. If root emergence does not occur during the first spring, the next opportunity for root emergence would be the third spring and epicotyl emergence would occur during the fourth spring after dispersal (Walck et al. 2005). Age to maturity, or flowering, is variable and has been recorded from 4 to 20 years depending on growing conditions (Case and Case 1997).
Terrestrial Habitats
Forest - Hardwood
Other Nations (1)
United StatesN4
ProvinceRankNative
WashingtonSNRYes
CaliforniaSNRYes
OregonSNRYes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentPervasive (71-100%)UnknownModerate (short-term)
7 - Natural system modificationsPervasive (71-100%)Moderate - slightHigh (continuing)
7.1 - Fire & fire suppressionPervasive (71-100%)Moderate - slightHigh (continuing)
7.1.1 - Increase in fire frequency/intensityPervasive (71-100%)Moderate - slightHigh (continuing)
8 - Invasive & other problematic species, genes & diseasesPervasive (71-100%)Moderate - slightHigh (continuing)
8.1 - Invasive non-native/alien species/diseasesPervasive (71-100%)Moderate - slightHigh (continuing)

Plant Characteristics
DurationPERENNIAL, Long-lived
Economic Value (Genus)Yes
Roadless Areas (41)
California (39)
AreaForestAcres
Bell QuinbyShasta-Trinity National Forest11,556
Big Butte ShinboneMendocino National Forest4,265
Black ButteMendocino National Forest15,461
Board CampSix Rivers National Forest4,527
Bonanza KingShasta-Trinity National Forest16,402
Box CampKlamath National Forest858
ChanchelullaShasta-Trinity National Forest3,915
ChinquapinShasta-Trinity National Forest22,040
Cow CreekShasta-Trinity National Forest22,627
Cow CreekSix Rivers National Forest1,271
East ForkShasta-Trinity National Forest6,201
East GirardShasta-Trinity National Forest27,894
GriderKlamath National Forest10,647
GrindstoneMendocino National Forest26,031
IshiLassen National Forest21,805
JohnsonKlamath National Forest10,652
KangarooKlamath National Forest40,617
Lake EleanorShasta-Trinity National Forest397
Little French CShasta-Trinity National Forest11,529
Middle ForkPlumas National Forest29,278
Murphy GladeShasta-Trinity National Forest1,015
North Fork American RiverTahoe National Forest38,495
Orleans Mtn.Klamath National Forest49,090
Orleans Mtn. BSix Rivers National Forest17,183
Orleans Mtn. CSix Rivers National Forest15,589
PantherShasta-Trinity National Forest12,016
PattisonShasta-Trinity National Forest29,299
Pilot CreekSix Rivers National Forest9,192
PyramidEldorado National Forest24,347
RussianKlamath National Forest21,771
Salt GulchShasta-Trinity National Forest6,511
ShacklefordKlamath National Forest1,679
SiskiyouKlamath National Forest54,039
Snow MountainMendocino National Forest14,457
SoliderSix Rivers National Forest14,918
South ForkShasta-Trinity National Forest16,786
ThatcherMendocino National Forest16,652
UnderwoodSix Rivers National Forest6,591
Weaver BallyShasta-Trinity National Forest829
Oregon (2)
AreaForestAcres
North KalmiopsisSiskiyou National Forests91,560
South KalmiopsisSiskiyou National Forests104,477
References (18)
  1. Bartuszevige, A.M., and B.A. Endress. 2008. Do ungulates facilitate native and exotic plant spread? Seed dispersal by cattle, elk and deer in northeastern Oregon. Journal of Arid Environments 72: 904-913.
  2. Case, F.W. and R.B. Case. 1997. Trilliums. Timber Press, Portland Oregon.
  3. Chafin, L. G. 2010d. Species account for <i>Trillium persistens </i>for Georgia Department of Natural Resources. Online. Available: georgiawildlife.com/sites/default/files/uploads/wildlife/nongame/pdf/accounts/plants/trillium_persistens.pdf.
  4. Chauhan, H., A. Bisht, I. Bhatt, A. Bhatt, and D. Gallacher. 2019. <i>Trillium </i>- toward sustainable utilization of a biologically distinct genus valued for traditional medicine. The Botanical Review 85(3): 252-272.
  5. <div>Meyers, S.C., T. Jaster, K. E. Mitchell, and L.K. Hardison, editors. 2015. Flora of Oregon. Volume 1: Pteridophytes, Gymnosperms, and Monocots. Botanical Research Institute of Texas, Fort Worth, TX. 608 pp.</div>
  6. Fernald, M.L., and A.C. Kinsey. 1943. Edible Wild Plants of Eastern North America. Idlewild Press, Cornwall-on-Hudson, NY. xiv+452 pp.
  7. Flora of North America Editorial Committee (FNA). 2002a. Flora of North America north of Mexico. Vol. 26. Magnoliophyta: Liliidae: Liliales and Orchidales. Oxford Univ. Press, New York. xxvi + 723 pp.
  8. Hitchcock, C.L., and A. Cronquist. 2018. Flora of the Pacific Northwest: An illustrated manual. Edited by D.E. Giblin, B. S. Legler, P.F. Zika, and R. G. Olmstead. 2nd edition. Univ. Washington Press, Seattle. xiii + 822 pp.
  9. Kartesz, J.T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. 2nd edition. 2 vols. Timber Press, Portland, OR.
  10. Klest, S.M. 2002. Propagation Protocol for Western Trilliums. Native Plants Journal 3(1):22-23.
  11. Leege, L. M., J. S. Thompson, D.J. Parris. 2010. The Responses of Rare and Common Trilliums (<i>Trillium reliquum</i>, <i>T. cuneatum</i>, and <i>T. maculatum</i>) to Deer Herbivory and Invasive Honeysuckle Removal. Castanea 75(4): 433-443.
  12. Lewis, Walter H., and Memory P.F. Elvin-Lewis. 1977. Medical Botany: Plants Affecting Man's Health. John Wiley and Sons, New York, New York. 515 p.
  13. Ohara, M. 1989. Life history evolution in the genus Trillium. Plant Species Biology 4:1-28.
  14. Rahman, S., M. Ismail, M. Khurram, I. Ullah, F. Rabbi, and M. Iriti. 2017. Bioactive steroids and saponins of the genus <i>Trillium. </i> Molecules 22(12): 2156.
  15. Trillium workshop group. 2019. , L. L. Gaddy, A. Floden, A. Frances, A. Highland, D. Leaman, T. Littlefield, C. Meredith, S. O'Bryan, L. Oliver, E. Schilling, A. Schotz, A. Walker, K. Wayman. Status assessment workshop at Mt. Cuba Center, Oct. 21-23, 2019.
  16. Vellend, M., J. Myers, S. Gardescu, and P. Marks. 2003. Dispersal of <i>Trillium</i> seeds by deer: Implications for long-distance migration of forest herbs. Ecology 84(4):1067-1072.
  17. Walck, J.L., J.M. Baskin, C.C. Baskin, and S.N. Hidayati. Defining transient and persistent seed banks in species with pronounced seasonal dormancy and germination patterns. Seed Science Research 15: 189-196. DOI: 10.1079/SSR2005209
  18. Zettler, J. A., T. P. Spira, and A. A. Craig. 2001. Yellow Jackets (<i>Vespula</i> spp.) Disperse <i>Trillium</i> (spp.) Seeds in Eastern North America. American Midland Naturalist 146(2):444-446.