Description
A medium-sized cormorant; males slightly larger than females. Adult basic or post-breeding plumage is blackish with glossy purple, bronze and green hues over the body and single-crested head, with dull reddish facial skin and yellowish bill with black tip and culmen (Causey 2002). Alternate or pre-breeding plumage is similar but with a striking white patch on the flank, brighter red facial skin and the presence of two head crests and white plumes trailing off the neck and/or breast area. Chicks hatch naked and are soon covered with brownish down, and juvenile birds are almost uniformly dark brown (plumage and bill) with feathered facial area, and develop naked forehead when reproductively mature (about 3 years) (Johnsgard 1993). Closely resembles the Pelagic Cormorant, which overlaps in range, but is larger with generally bigger proportions (Causey 2002).
Habitat
Closely associated with rock-bottom coastlines of North Pacific marine islands and isolated areas of mainland Alaska, Kamchatka and Japan; often close to shore in water less than 200 m deep. Nests on steep, relatively inaccessible slopes, on ledges averaging about 40 cm wide (Johnsgard 1993), including rocky sea islands alongside gulls, murres, and auklets. Roosts on offshore rocks or protected cliff outcroppings and forages in water near shore (< 20 km) (Causey 2002).
Ecology
Species is exclusively marine, coming to land only to breed or roost (Causey 2002). Predators include foxes, gulls, corvids, and raptors (e.g., bald eagle); glaucous-winged gull and ravens may be serious egg predators in some areas; human disturbance resulting in predation by American crows or other predators; attempted predation by river otter was a significant decimating factor on Kodiak Island (see Johnsgard 1993).
Reproduction
Eggs are laid in May-June in Alaska; mean laying date in mid- or late May but time of nesting varies between years. Clutch size usually is 2-4. Incubation, by both sexes, lasts about 31-34 days (average). Young fledge at average age estimated at about 7 to 8+ weeks (beginning in mid-August or as late as early September in Alaska). Productivity and local abundance may vary greatly from year to year.