Ursus arctos

Linnaeus, 1758

Brown Bear

G4Apparently Secure Found in 120 roadless areas NatureServe Explorer →
G4Apparently SecureGlobal Rank
Least concernIUCN
PS:E,T,XNESA Status
HighThreat Impact
Ursus arctos. Photo by Константин Шатилов, via iNaturalist.
Константин Шатилов, CC BY 4.0
Ursus arctos. Photo by Михаил Голомысов, via iNaturalist.
Михаил Голомысов, CC BY 4.0
Ursus arctos. Photo by Simon Rolph, via iNaturalist.
Simon Rolph, CC BY 4.0
Ursus arctos. Photo by Simon Rolph, via iNaturalist.
Simon Rolph, CC BY 4.0
Ursus arctos. Photo by 松井 淳, via iNaturalist.
松井 淳, CC0 1.0
Ursus arctos. Photo by Matt Muir, via iNaturalist.
Matt Muir, CC BY 4.0
Grizzly bear (Ursus arctos horribilis). Photo by U.S. Fish & Wildlife Service, Public Domain (U.S. Government Work), via ECOS.
U.S. Fish & Wildlife Service, https://www.usa.gov/government-works
Identity
Unique IDELEMENT_GLOBAL.2.102187
Element CodeAMAJB01020
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
CITESAppendix II
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassMammalia
OrderCarnivora
FamilyUrsidae
GenusUrsus
USESAPS:E,T,XN
COSEWICX,SC
Other Common Names
brown bear (EN) Grizzly Bear (EN) Mexican Grizzly Bear (EN) Ours brun (FR) Ours grizzli (FR)
Concept Reference
Wilson, D. E., and D. M. Reeder (editors). 1993. Mammal species of the world: a taxonomic and geographic reference. Second edition. Smithsonian Institution Press, Washington, DC. xviii + 1206 pp. Available online at: http://www.nmnh.si.edu/msw/.
Taxonomic Comments
Genetic studies of brown bears indicate that the traditional morphology-based taxonomy of brown bears is highly discordant with bear phylogeny as indicated by geographic patterns of mtDNA variation. Based on recent and permafrost-preserved Pleistocene material, there is no genetic (mtDNA) support for the validity of any of the commonly recognized North American subspecies (e.g., horribilis, middendorffi), and North American brown bears do not represent a distinct lineage with respect to brown bears in Northern Asia and Europe (Waits et al. 1998, Leonard et al. 2000, Barnes et al. 2002). If a subspecific name is to be applied to North American brown bears, it should be Ursus arctos arctos, a taxon whose range encompasses both North America and parts of Eurasia. This name has been adopted for North American brown bears by ITIS (http://www.itis.usda.gov/index.html), which lists U. a. horribilis and U. a. nelsoni as invalid because they are junior synonyms of U. a. arctos.

There are six recovery ecosystems for grizzly bears in the lower-48 US states: The Greater Yellowstone (GYE), the Northern Continental Divide (NCDE), the North Cascades (NCE), the Selkirks (SE), the Cabinet-Yaak (CYE), and the Bitterroot (BE).
Conservation Status
Rank Method Rank calculation - Biotics v2
Review Date2022-07-25
Change Date1996-11-18
Edition Date2022-07-25
Edition AuthorsCannings, S. G., and G. Hammerson; rev. B. Young (2022)
Threat ImpactHigh
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences81 to >300
Rank Reasons
This species has a very large Holarctic range where it has declined in Europe and North America and faces numerous threats. However, large portions of the range are in remote areas where there is little evidence of population declines. Intensive management in Europe and North America have helped stabilize and recover some subpopulations.
Range Extent Comments
Formerly throughout western North America, north from northern Mexico; northwestern Africa, all of the Palearctic from western Europe, Near and Middle East through the northern Himalayas to western and northern China and Chukot (Russia) and Hokkaido (Japan) (Wozencraft, in Wilson and Reeder 1993); see Pasitschniak-Arts (1993) for additional details. In North America, present range includes Alaska, northern and western Canada, northern Continental Divide in Montana, Cabinet/Yaak mountains in Montana/Idaho, and Yellowstone area, Wyoming/Montana/Idaho, USA. Common only in Alaska, parts of the Yukon, northern and coastal British Columbia, and portions of the northern Rocky Mountains. In Europe, apart from northern Europe, distribution has shrunk to a few isolated populations in the Pyrenees, the Apenines, the Alps, the Balkan Peninsula, and the Carpathians (Hartl and Hell 1994), although reintroductions have bolstered subpopulations in some areas (Barba et al. 2010).
Occurrences Comments
This species is known from hundreds of distinct localities, which approximate occurrences, with records from the period 2000-2022 (GBIF 2022).
Threat Impact Comments
Historical decline due to habitat loss and fragmentation and killing by humans. Primary threats are habitat alienation, alteration, and loss; increased access to wilderness; and hunting (both legal and illegal). Increased access increases human-bear contacts, some of which result in destruction of bears. Alien species threaten food resources in some areas; in Montana, white pine blister rust has killed whitebark pines (seeds serve as food for bears) and knapweed have displaced native plants that serve as foods for bears and their prey. See Horejsi (1989) for a discussion of land-use threats (petroleum and natural gas development, grazing by domestic stock) and excessive bear mortality in southwestern Alberta. See also Matthews and Moseley (1990) for discussion of threats. Several large mines in Montana, if approved, may pose a threat (USFWS 1999). Forestry, mining, recreation, and road building also affect habitat in British Columbia where the two portions of this distinct population segment are connected (USFWS 1999, Proctor et al. 2022).
Ecology & Habitat

Description

Color ranges from pale yellowish to dark brown; usually white tips on the hairs, especially on the back, resulting in a frosted or grizzled effect; facial profile concave; claws on front feet of adults about 4 inches long and curved; noticeable hump above shoulders; head and body of adults about 6-8 feet, height at shoulders 3-4.5 feet (Burt and Grossenheider 1964).

Diagnostic Characteristics

Differs from black bear in being larger as an adult and by having a hump above the shoulders and a concave (rather than straight or convex) facial profile.

Habitat

Now found mostly in arctic tundra, alpine tundra, and subalpine mountain forests. Once found in a wide variety of habitats including: open prairie, brushlands, riparian woodlands, and semidesert scrub. Ranges widely at the landscape level. Most populations require huge areas of suitable habitat. Common only where food is abundant and concentrated (e.g., salmon runs, caribou calving grounds). Typically digs own hibernation den, usually on steep northern slope where snow accumulates. See LeFranc et al. (1987).

Young are born in den in cave, crevice, hollow tree, hollow dug under rock, or similar site. Use of summit or ridge for mating (in May-June) reported for Banff National Park, Alberta, but not elsewhere (Hamer and Herrero 1990). In the Northwest Territories, Canada, all dens were on well-drained slopes; the majority of dens faced south (25), followed by west (13), east (10), and north (8); most dens were constructed under cover of tall shrubs (Betula glandulosa and Salix), the root structures of which supported ceilings of dens; esker habitat was selected more than expected by chance (McLoughlin et al. 2002).

In Spain, remnant deciduous forests and upland creek drainages were prime feeding areas (Clevenger et al. 1992).

Ecology

May congregate in areas with abundant food; otherwise solitary except when breeding or caring for young. Density estimates range from 1/1.5-4 sq km (Kodiak Island) to 1/50 sq km (Yellowstone) to 0.6-7.9/1000 sq km (Norway).

In the Yellowstone region, lack of berries and large fluctuations in the size of pine seed crops were major factors limiting bear density (Mattson et al. 1991).

In British Columbia-Montana, survivorship of adult and subadult females was the most important variable in estimating population trend.

Reproduction

Breeds in late spring and early summer. Implantation is delayed; gestation lasts about 184 days. Litter size is 1-4 (average 2). Young are born in winter, remain with mother usually the first two winters. Breeding interval generally is 2-4 years. In North America, first parturition occurs at 5-6 years in the south, 6-9 years in the north. A few live as long as 20-25 years. Long life span, late sexual maturity, protracted reproductive cycles.
Terrestrial Habitats
Forest - HardwoodForest - ConiferForest - MixedWoodland - HardwoodWoodland - ConiferWoodland - MixedShrubland/chaparralGrassland/herbaceousAlpineTundra
Palustrine Habitats
Riparian
Other Nations (2)
CanadaN3
ProvinceRankNative
British ColumbiaS3Yes
AlbertaS3Yes
Northwest TerritoriesS3Yes
SaskatchewanSXYes
ManitobaSXYes
NunavutS3Yes
LabradorSXYes
Yukon TerritoryS3Yes
QuebecSXYes
United StatesN3
ProvinceRankNative
North DakotaSXYes
UtahSXYes
AlaskaS4Yes
New MexicoSXYes
ArizonaSXYes
MontanaS3Yes
WashingtonS1Yes
ColoradoSXYes
NevadaSXYes
MinnesotaSXYes
South DakotaSXYes
OklahomaSXYes
NebraskaSXYes
CaliforniaSXYes
OregonSXYes
TexasSXYes
WyomingS1Yes
KansasSXYes
IdahoSNRYes
Navajo NationSXYes
Threat Assessments
ThreatScopeSeverityTiming
1 - Residential & commercial developmentSmall (1-10%)Extreme - seriousHigh (continuing)
1.1 - Housing & urban areasSmall (1-10%)Extreme - seriousHigh (continuing)
1.2 - Commercial & industrial areasSmall (1-10%)Serious or 31-70% pop. declineHigh (continuing)
2 - Agriculture & aquacultureSmall (1-10%)Serious or 31-70% pop. declineHigh (continuing)
2.1 - Annual & perennial non-timber cropsSmall (1-10%)Serious or 31-70% pop. declineHigh (continuing)
2.2 - Wood & pulp plantationsSmall (1-10%)Serious or 31-70% pop. declineHigh (continuing)
3 - Energy production & miningSmall (1-10%)Serious or 31-70% pop. decline
3.2 - Mining & quarryingSmall (1-10%)Serious or 31-70% pop. decline
4 - Transportation & service corridorsSmall (1-10%)Serious - moderateHigh (continuing)
4.1 - Roads & railroadsSmall (1-10%)Serious - moderateHigh (continuing)
5 - Biological resource useLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)
5.1 - Hunting & collecting terrestrial animalsLarge (31-70%)Moderate or 11-30% pop. declineHigh (continuing)
6 - Human intrusions & disturbanceSmall (1-10%)Slight or 1-10% pop. decline
6.1 - Recreational activitiesSmall (1-10%)Slight or 1-10% pop. decline

Roadless Areas (120)
Alaska (60)
AreaForestAcres
AnanTongass National Forest36,666
Bering LakeChugach National Forest965,076
Boston BarChugach National Forest53,617
Brabazon AdditionTongass National Forest498,819
CastleTongass National Forest49,289
ChichagofTongass National Forest555,858
Chilkat-West Lynn CanalTongass National Forest199,772
Chugach-12Chugach National Forest8,116
Chugach-13Chugach National Forest13,337
ClevelandTongass National Forest189,447
College FiordChugach National Forest1,130,818
Copper River WetlandsChugach National Forest85,972
Douglas IslandTongass National Forest28,065
Fidalgo-GravinaChugach National Forest257,968
Five MileTongass National Forest19,459
Freshwater BayTongass National Forest44,933
Game CreekTongass National Forest54,469
Greens CreekTongass National Forest27,177
HardingTongass National Forest174,349
Hoonah SoundTongass National Forest79,764
HyderTongass National Forest121,723
Johnson PassChugach National Forest152,508
Juneau UrbanTongass National Forest101,581
Juneau-Skagway IcefieldTongass National Forest1,187,268
Kenai LakeChugach National Forest213,172
Kenai MountainsChugach National Forest306,600
MadanTongass National Forest68,553
Mansfield PeninsulaTongass National Forest54,991
Middle KruzofTongass National Forest14,724
MontagueChugach National Forest204,875
MosmanTongass National Forest53,492
Nellie JuanChugach National Forest713,697
North BaranofTongass National Forest314,089
North ClevelandTongass National Forest105,288
North KruzofTongass National Forest33,146
North KupreanofTongass National Forest114,660
North RevillaTongass National Forest215,430
North WrangellTongass National Forest8,091
Pavlof-East PointTongass National Forest5,399
Point AugustaTongass National Forest15,482
Point CravenTongass National Forest10,906
Port AlexanderTongass National Forest120,681
Prince William Sound Is.Chugach National Forest118,698
RedoubtTongass National Forest68,347
ResurrectionChugach National Forest224,615
RhineTongass National Forest23,010
Roaded DonutChugach National Forest968
Sheridan GlacierChugach National Forest224,683
Sitka SoundTongass National Forest13,459
Sitka UrbanTongass National Forest112,003
South KruzofTongass National Forest55,193
South KupreanofTongass National Forest216,816
SullivanTongass National Forest67,323
Taku-SnettishamTongass National Forest664,928
Tenakee RidgeTongass National Forest20,527
Twenty MileChugach National Forest198,775
Upper SitukTongass National Forest16,789
WhitestoneTongass National Forest5,638
Windham-Port HoughtonTongass National Forest161,952
Yakutat ForelandsTongass National Forest323,648
Idaho (3)
AreaForestAcres
HoodooNez Perce-Clearwater National Forest153,868
Italian PeakCaribou-Targhee National Forest141,158
PalisadesCaribou-Targhee National Forest122,002
Montana (28)
AreaForestAcres
Bear - Marshall - Scapegoat - SwanLewis and Clark National Forest344,022
Bear - Marshall - Scapegoat - SwanLolo National Forest118,485
BeartoothGallatin National Forest5,285
Bmss Ra 1485Flathead National Forest334,275
Cabin Creek Wildlife Management Area OcdGallatin National Forest35,048
Cabinet Face East #671Kootenai National Forest50,326
Coal Ridge Ra 1127Flathead National Forest15,429
Deadhorse Ridge Ra 1128Flathead National Forest23,648
East PioneerBeaverhead-Deerlodge National Forest145,082
Freezeout MountainBeaverhead-Deerlodge National Forest97,305
Gallatin FringeGallatin National Forest51,571
Garfield MountainBeaverhead-Deerlodge National Forest41,891
Hyalite - Porcupine - Buffalo Horn Wilderness Study AreaGallatin National Forest143,991
Line Creek PlateauCuster National Forest24,825
LionheadGallatin National Forest33,549
MadisonGallatin National Forest127,859
Middle Fork Judith WsaLewis and Clark National Forest81,131
Mt. Gmt Area HCuster National Forest1,335
North AbsarokaGallatin National Forest159,075
North AbsarokaCuster National Forest21,063
North Big HoleBeaverhead-Deerlodge National Forest52,227
North CarpBeaverhead-Deerlodge National Forest6,600
Red Lodge Creek HellroaringCuster National Forest17,210
ReefGallatin National Forest2,500
SapphireBitterroot National Forest43,303
SapphiresBeaverhead-Deerlodge National Forest66,619
Snowcrest MountainBeaverhead-Deerlodge National Forest97,649
West PioneerBeaverhead-Deerlodge National Forest248,631
Washington (3)
AreaForestAcres
Grassy TopIdaho Panhandle National Forests13,485
Grassy TopColville National Forest10,302
Harvey CreekColville National Forest10,140
Wyoming (26)
AreaForestAcres
Beartooth Proposed WildernessShoshone National Forest16,837
Bench MarkShoshone National Forest5,940
Franc's PeakShoshone National Forest68,561
Grayback RidgeBridger-Teton National Forest295,113
Gros Ventre MountainsBridger-Teton National Forest106,418
Headwaters Sunlight CreekShoshone National Forest6,016
Little Sheep MountainBridger-Teton National Forest14,192
Mosquito Lake - Seven LakesBridger-Teton National Forest51,950
Pacific Creek - Blackrock CreekBridger-Teton National Forest24,658
PalisadesTarghee National Forest1,121
Phillips RidgeBridger-Teton National Forest10,108
ReefShoshone National Forest16,817
Sheridan PassShoshone National Forest8,394
Sleeping GiantShoshone National Forest5,176
South Beartooth HighwayShoshone National Forest105,570
South ForkShoshone National Forest64,903
Spread Creek - Gros Ventre RiverBridger-Teton National Forest166,097
Sulphur CreekShoshone National Forest30,221
Teton Corridor TrailheadsBridger-Teton National Forest286
Togwotee PassShoshone National Forest7,344
Trout CreekShoshone National Forest44,034
Wapiti Valley EastShoshone National Forest449
Wapiti Valley NorthShoshone National Forest18,345
Wapiti Valley SouthShoshone National Forest43,517
West Slope TetonsTarghee National Forest47,448
Windy MountainShoshone National Forest31,283
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