Chrysemys picta

(Schneider, 1783)

Painted Turtle

G5Secure Found in 80 roadless areas NatureServe Explorer →
G5SecureGlobal Rank
Least concernIUCN
LowThreat Impact
Identity
Unique IDELEMENT_GLOBAL.2.637723
Element CodeARAAD01010
Record TypeSPECIES
ClassificationSpecies
Classification StatusStandard
Name CategoryVertebrate Animal
IUCNLeast concern
Endemicoccurs (regularly, as a native taxon) in multiple nations
KingdomAnimalia
PhylumCraniata
ClassChelonia
OrderTestudines
FamilyEmydidae
GenusChrysemys
COSEWICPS:T,SC,NAR
Other Common Names
northern painted turtle (EN) Tortue peinte (FR)
Concept Reference
Starkey, D. E., H. B. Shaffer, R. L. Burke, M.R.J. Forstner, J. B. Iverson, F. J. Janzen, A.G.J. Rhodin, and G. R. Ultsch. 2003. Molecular systematics, phylogeography, and the effects of Pleistocene glaciation in the painted turtle (Chrysemys picta) complex. Evolution 57:119-128.
Taxonomic Comments
Includes three subspecies: C. p. bellii, C. p. marginata, and C. p. picta. Based on molecular data, Starkey et al. (2003) recognized C. dorsalis as a distinct species. This was adopted by Crother et al. (2008, 2017) and the Turtle Taxonomy Working Group (2012, 2021). Some herpetologists, citing apparent intergrades in western Kentucky, southern Illinois, and southeastern Missouri, still regard it as conspecific with C. picta. Additional genetic work by Jensen et al. (2014) and Jensen et al. (2015) supported Starkey et al. (2003). However, according to Crother (2017), these studies also questioned the recognition of the remaining subspecies on genetic grounds, but did not take a position on their abandonment. In contrast, Ernst et al. (2006) reexamined color patterns and dorsal scute alignment in Chrysemys and identified intermediate specimens between C. dorsalis and C. p. marginata and C. p. bellii. Although the evidence for species recognition of dorsalis is primarily mitochondrial, Crother (2017) continue to recognize it as a full species pending further genetic sampling.
Conservation Status
Rank Method Rank calculation - Biotics v2
Review Date2024-02-04
Change Date1996-10-21
Edition Date2024-02-04
Edition AuthorsGundy, R. L.
Threat ImpactLow
Range Extent>2,500,000 square km (greater than 1,000,000 square miles)
Number of Occurrences> 300
Rank Reasons
This species is widely distributed across North America and abundant throughout most of its range. It is highly adaptable to altered and man-made aquatic habitats. There are no major range-wide declines reported. However, some western populations are rare and declining. There are no major range-wide threats, although there are localized threats including habitat, loss, predation, severe flooding, pollution, and drought.
Range Extent Comments
This species is the only turtle species that spans the width of North America from coast to coast (Ernst and Lovich 2009). It ranges from New Brunswick and Nova Scotia, Canada across southern Canada to Vancouver Island, British Columbia, Canada (Ernst and Lovich 2009, Barela and Olson 2014, COSEWIC 2016). It is found in most states in the eastern and central United States. The range extends farthest south in the east where it reaches the Gulf Coast in Louisiana, United States (Ernst and Lovich 2009). There are three subspecies. Chrysemys picta picta occurs along the Coastal Plain and Appalachians east of the Great Lakes, C. p. marginata occurs in the Great Lakes Region east of the Mississippi River, and C. p. bellii occurs west of the Mississippi River (Starkey et al. 2003, Ernst and Lovich 2009, Ernst et al. 2014). Using a combination of records from Global Biodiversity Information Facility (GBIF 2024) and iNaturalist (2024), GeoCat (2024) estimated range extent to be about 7,250,000 km².
Occurrences Comments
This species has many hundreds of occurrences throughout its range (Ernst and Lovich 2009, Barela and Olson 2014, COSEWIC 2016).
Threat Impact Comments
There are no major range-wide threats affecting this species. There are a number of threats operating at local scales including habitat, loss, predation, severe flooding, pollution, and drought (Ernst and Lovich 2009). Development of uplands surrounding occupied ponds may cause long-term declines that are not detectable for many years (Eskew et al. 2010, Capkun-Huot et al. 2021). Roads are generally suspected of causing a high degree of mortality due to vehicle strikes and a lack of connectivity between ponds, but there is a growing body of evidence documenting a remarkable adaptability and resilience to roads and some development (Ernst and Lovich 2009, Laporte et al. 2012, Dorland et al. 2014, Capkun-Huot et al. 2021). Western populations are threatened by habitat loss due to development and agriculture (Barela and Olson 2014, COSEWIC 2016).
Ecology & Habitat

Description

Shell hard, somewhat flattened; bright yellow lines on head and limbs; lower shell orange or reddish, with conspicuous dark markings in juveniles; upper shell often with narrow yellow lines (reduced in larger individuals), less often with dark reticulation; upper jaw notched at tip; maximum upper shell length of female about 25 cm, rarely more than 21 cm (males are much smaller). Mature male: vent located beyond rear edge of upper shell with tail extended; fingernails very long; lower shell not concave. Mature female: vent at or inside rear edge of upper shell with tail extended; fingernails not especially long. Juvenile: as in adult female, with a deep crease across abdominal shields of lower shell. Hatchling: upper shell keeled; lower shell vivid orange/red, with a central dark figure having a sinuous outer edge; upper shell length 19-28 mm (average 23-24 mm). Eggs: elliptical, whitish, smooth surface with small pores; shell initially flexible, then becomes more rigid, 21-36 mm x 15-21 mm (mostly about 29-33 x 18-19 mm). Source: Hammerson (1999).

Habitat

This species lives in slow-moving, shallow waters with soft bottoms such as rivers, streams, marshes, swamps, ponds, lakes, and reservoirs (Ernst and Lovich 2009). Aquatic vegetation and abundant basking sites are important habitat components (Ernst and Lovich 2009). They may colonize seasonally flooded areas near permanent water (Ernst and Lovich 2009). Nesting occurs in soft soil up to several hundred meters from water (Christens and Bider 1987, Congdon and Gatten 1989, Ernst and Lovich 2009). Some hatchlings remain in the nest in winter and emerge in spring (Packard and Packard 1995, Ernst and Lovich 2009). Adults estivate in water (Ernst and Lovich 2009).

Ecology

Home ranges vary drastically. Lakes, ponds, and marsh populations generally have the smallest home ranges and the largest home ranges generally occur in river or creek systems (Ernst and Lovich 2009). Population densities vary widely with the greatest densities, up to several hundred individuals per hectare, occurring in lakes, ponds, and marshes, while river or creek systems generally have the lowest population densities (Ernst and Lovich 2009). Survivorship of eggs and hatchlings is generally below 20% (Christens and Bider 1987, Lindeman 1991, Ernst and Lovich 2009). Survivorship rates increase with age, with adults generally having survivorship rates above 50% and many reports are above 90% (Mitchell 1988, Frazer et al. 1991, Iverson and Smith 1993, Ernst and Lovich 2009). See Iverson (1991) for a compilation of survivorship data on eggs and juveniles. In Nebraska, some females apparently survived beyond 30 years (Iverson and Smith 1993).

Reproduction

Most nesting occurs between late May and early July, perhaps earlier in the south (Ernst and Lovich 2009). Females dig nests in soft soil in open areas up to several hundred meters from water (Christens and Bider 1987, Congdon and Gatten 1989, Ernst and Lovich 2009). Individual females can produce up to five clutches per season (Ernst and Lovich 2009). Clutch size ranges from 1-23 eggs, with the largest subspecies, C. p. bellii, producing the largest clutch sizes and the smallest, C. p. picta, producing the smallest clutch sizes (Ernst and Lovich 2009). Females reach sexual maturity in as little as 5 years in southern parts of its range and as much as 10 years in northern parts of its range (Ernst and Lovich 2009). Males mature at younger ages (Frazzer et al. 1993, Ernst and Lovich 2009). Hatchlings usually remain in the nest in winter, tolerating freezing temperatures, and emerge in spring (Ernst and Lovich 2009).
Palustrine Habitats
TEMPORARY POOLHERBACEOUS WETLANDSCRUB-SHRUB WETLANDFORESTED WETLANDRiparian
Other Nations (2)
United StatesN5
ProvinceRankNative
OhioSNRYes
ArizonaS1Yes
New JerseySNRYes
ArkansasS3Yes
IdahoS3Yes
IllinoisSNRYes
WashingtonS4Yes
MaineS5Yes
MichiganS5Yes
North CarolinaS5Yes
KansasS5Yes
UtahSNAYes
MarylandS5Yes
New YorkS5Yes
ConnecticutS5Yes
IowaS5Yes
South DakotaS5Yes
OregonS2Yes
New MexicoS4Yes
MinnesotaS5Yes
NebraskaS5Yes
KentuckySNRYes
MassachusettsS5Yes
New HampshireS5Yes
TennesseeS5Yes
WisconsinS5Yes
MontanaS5Yes
GeorgiaS5Yes
Rhode IslandS5Yes
ColoradoS5Yes
Navajo NationSNAYes
WyomingS4Yes
VermontS5Yes
VirginiaS5Yes
South CarolinaSNRYes
IndianaS4Yes
OklahomaS2Yes
NevadaSNANo
PennsylvaniaS5Yes
District of ColumbiaS5Yes
West VirginiaS5Yes
TexasS1Yes
MissouriS5Yes
North DakotaSNRYes
AlabamaSNRYes
DelawareS5Yes
CanadaN5
ProvinceRankNative
New BrunswickS4Yes
OntarioS4Yes
British ColumbiaS3Yes
Island of NewfoundlandSNANo
QuebecS4Yes
SaskatchewanS3Yes
Nova ScotiaS4Yes
AlbertaS2Yes
ManitobaS4Yes
Threat Assessments
ThreatScopeSeverityTiming
No known threats

Roadless Areas (80)
Georgia (1)
AreaForestAcres
Helton CreekChattahoochee National Forest2,348
Idaho (2)
AreaForestAcres
Bear CreekCaribou-Targhee National Forest118,582
HoodooNez Perce-Clearwater National Forest153,868
Illinois (1)
AreaForestAcres
Bay CreekShawnee National Forest120
Minnesota (5)
AreaForestAcres
Baker - Homer - Brule LakesSuperior National Forest6,712
Brule Lake - Eagle MountainSuperior National Forest12,380
Kawishiwi Lake To SawbillSuperior National Forest15,305
Little Indian SiouxSuperior National Forest996
Wood LakeSuperior National Forest596
Montana (21)
AreaForestAcres
Bear - Marshall - Scapegoat - SwanLewis and Clark National Forest344,022
Bear - Marshall - Scapegoat - SwanHelena National Forest51,360
Bear - Marshall - Scapegoat - SwanLolo National Forest118,485
Big Snowy Mountains WsaLewis and Clark National Forest88,003
Bmss Ra 1485Flathead National Forest334,275
BridgerGallatin National Forest45,059
Buckhorn Ridge (MT)Kootenai National Forest34,716
Cabinet Face East #671Kootenai National Forest50,326
Cube Iron - SilcoxLolo National Forest36,998
East Fork Elk #678Kootenai National Forest6,767
Freezeout MountainBeaverhead-Deerlodge National Forest97,305
Lazyman GulchHelena National Forest11,614
Lone Cliff West #674aKootenai National Forest5,315
Marston Face # 172Kootenai National Forest9,098
Mt. Gmt Area HCuster National Forest1,335
North AbsarokaCuster National Forest21,063
Ogden MountainHelena National Forest12,148
Scotchman Peaks (MT)Kootenai National Forest53,909
Selway - Bitterroot (01067)Bitterroot National Forest114,953
Tongue River BreaksCuster National Forest17,481
West Big HoleBeaverhead-Deerlodge National Forest133,563
New Hampshire (4)
AreaForestAcres
Carr MountainWhite Mountain National Forest17,110
PemigewassetWhite Mountain National Forest32,255
Sandwich RangeWhite Mountain National Forest16,797
Wild RiverWhite Mountain National Forest46,878
North Dakota (3)
AreaForestAcres
Bullion ButteDakota Prairie Grasslands19,877
MagpieDakota Prairie Grasslands21,281
Ponderosa PineDakota Prairie Grasslands7,471
Oregon (4)
AreaForestAcres
Castle RidgeWallowa-Whitman National Forest8,377
EagleMt. Hood National Forest16,841
HellholeUmatilla National Forest65,679
Hurricane CreekWallowa-Whitman National Forest1,606
South Dakota (1)
AreaForestAcres
Indian CreekBuffalo Gap National Grassland24,666
Tennessee (1)
AreaForestAcres
Sampson Mountain AdditionCherokee National Forest3,064
Texas (1)
AreaForestAcres
Big CreekNational Forests in Texas1,447
Utah (6)
AreaForestAcres
418024Uinta National Forest51,699
Burch CreekWasatch-Cache National Forest6,938
HogsbackWasatch-Cache National Forest7,936
Mt. Logan NorthWasatch-Cache National Forest18,930
Signal PeakFishlake National Forest30,889
WellsvilleWasatch-Cache National Forest1,717
Vermont (2)
AreaForestAcres
Bread LoafGreen Mountain and Finger Lakes National Forests1,768
Griffith Lake 09084Green Mountain and Finger Lakes National Forests1,833
Virginia (14)
AreaForestAcres
Bear CreekJefferson National Forest18,274
Beards MountainGeorge Washington National Forest7,505
Broad RunJefferson National Forest10,971
Brush MountainJefferson National Forest6,002
Garden MountainJefferson National Forest3,960
Kelley MountainGeorge Washington National Forest7,590
Mill MountainGeorge Washington National Forest10,840
Mountain Lake Addition AJefferson National Forest1,469
North MountainJefferson National Forest8,377
Northern MassanuttenGeorge Washington National Forest9,444
Oliver MountainGeorge Washington National Forest13,090
Seng MountainJefferson National Forest6,428
Shawvers Run AdditionJefferson National Forest1,927
Southern MassanuttenGeorge Washington National Forest11,985
Washington (11)
AreaForestAcres
Black CanyonOkanogan National Forest9,681
Black CanyonWenatchee National Forest176
EntiatWenatchee National Forest72,617
Granite MountainOkanogan National Forest27,428
Grassy TopColville National Forest10,302
Mt. Baker Noisy - DiobsudMt Baker-Snoqualmie National Forest56,039
Rock CreekWenatchee National Forest32,239
Salmo - Priest BColville National Forest11,869
SawtoothOkanogan National Forest122,194
Silver StarGifford Pinchot National Forest7,779
TaneumWenatchee National Forest26,140
West Virginia (2)
AreaForestAcres
Canaan LoopMonongahela National Forest7,867
Mcgowan MountainMonongahela National Forest10,504
Wisconsin (1)
AreaForestAcres
09166 - East TorchChequamegon-Nicolet National Forest4,647
References (48)
  1. Barela, K. L., and D. H. Olson. 2014. Mapping the western pond turtle (<i>Actinemys marmorata</i>) and painted turtle (<i>Chrysemys picta</i>) in western North America. Northwestern Naturalist 95(1): 1-12.
  2. Capkun-Huot, C., V. K. Fyson, and G. Blouin-Demers. 2021. Landscape composition predicts the local abundance of painted turtles (<i>Chrysemys picta</i>). Herpetology Notes 14: 215-223.
  3. Christens, E., and J. R. Bider. 1987. Nesting activity and hatching success of the painted turtle (<i>Chrysemys picta marginata</i>) in southwestern Quebec. Herpetologica 43:55-65
  4. Christiansen, J. L., and E. O. Moll. 1973. Latitudinal reproductive variation in a single subspecies of painted turtle, <i>Chrysemys picta bellii</i>. Herpetologica 29:152-163.
  5. Conant, R. and J. T. Collins. 1991. A field guide to reptiles and amphibians: eastern and central North America. Third edition. Houghton Mifflin Co., Boston, Massachusetts. 450 pp.
  6. Congdon, J. D., and R. E. Gatten, Jr. 1989. Movements and energetics of nesting <i>Chrysemys picta</i>. Herpetologica 45:94-100.
  7. COSEWIC. 2016. COSEWIC assessment and status report on the Western Painted Turtle <i>Chrysemys picta bellii</i>, Pacific Coast population, Intermountain – Rocky Mountain population and Prairie/Western Boreal – Canadian Shield population,in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa. xxi + 95 pp. Available at https://www.canada.ca/en/environment-climate-change/services/species-risk-public-registry/cosewic-assessments-status-reports/western-painted-turtle-2016.html#_exesum.
  8. Crother, B. I. (editor). 2008. Scientific and standard English names of amphibians and reptiles of North America north of Mexico, with comments regarding confidence in our understanding. Sixth edition. Society for the Study of Amphibians and Reptiles Herpetological Circular 37:1-84. Online with updates at: http://www.ssarherps.org/pages/comm_names/Index.php
  9. Crother, B. I. (editor). 2017. Scientific and standard English names of amphibians and reptiles of North America north of Mexico, with comments regarding confidence in our understanding. 8th edition. SSAR Herpetological Circular 43:1-104. [Updates in SSAR North American Species Names Database at: https://ssarherps.org/cndb]
  10. DeGraaf, R. M., and D. D. Rudis. 1983a. Amphibians and reptiles of New England. Habitats and natural history. Univ. Massachusetts Press. vii + 83 pp.
  11. Dorland, A., T. Rytwinski, and L. Fahrig. 2014. Do roads reduce painted turtle (<i>Chrysemys picta</i>) populations? PLoSONE9(5): e98414.doi:10.1371/journal. pone.0098414
  12. Ernst, C. H. 1971. Chrysemys picta. Cat. Am. Amph. Rep. 106.1-106.4.
  13. Ernst, C. H. 1988. <i>Chrysemys</i>. Cat. Am. Amph. Rep. 438: 1-8.
  14. Ernst, C.H., A.F. Laemmerzahl, and T.R. Creque. 2006. A review of morphological and pattern variation in the painted turtle, <i>Chrysemys picta</i>, in Missouri, USA, with an alternate hypothesis of the origin of <i>Chrysemys picta marginata</i>. Herpetological bulletin 95:6-15.
  15. Ernst. C. H., and J. E. Lovich. 2009. Turtles of the United States and Canada. Second edition, revised and updated. Johns Hopkins University Press, Baltimore, Maryland. xii + 827 pp.
  16. Ernst, C. H., and R. W. Barbour. 1972. Turtles of the United States. Univ. Press of Kentucky, Lexington. x + 347 pp.
  17. Ernst, C. H., J. M. Orr, A. F. Laemmerzahl, and T. R. Creque. 2014. Variation and zoogeography of the turtle <i>Chrysemys picta</i> in Virginia, USA. The Herpetological Bulletin 130: 9-15.
  18. Ernst, C. H., R. W. Barbour, and J. E. Lovich. 1994. Turtles of the United States and Canada. Smithsonian Institution Press, Washington, D.C. xxxviii + 578 pp.
  19. Eskew, E. A., S. J. Price, and M. E. Dorcas. 2010. Survivorship and population densities of painted turtles (<i>Chrysemys picta</i>) in recently modified suburban landscapes. Chelonian Conservation and Biology 9(2): 244-249.
  20. Frazer, N. B., J. W. Gibbons, and J. L. Greene. 1991. Growth, survivorship and longevity of painted turtles Chrysemys picta in a southwestern Michigan marsh. Am. Midl. Nat. 125:245-258.
  21. Geospatial Conservation Assessment Tool (GeoCAT). Online. Available: <a href="http://geocat.kew.org/">https://geocat.iucnredlist.org/</a> (Accessed 2024)
  22. Gibbons, J.W. 1968a. Population structure and survivorship in the painted turtle, Chrysemys picta. Copeia 1968(2): 260-268.
  23. Gibbons, J. W. 1968c. Reproductive potential, activity and cycles in the painted turtle, <i>Chrysemys picta</i>. Ecology 49:399-409.
  24. Global Biodiversity Information Facility (GBIF). 2024. Global Biodiversity Information Facility (GBIF) data portal. Online. Available: https://www.gbif.org/ (accessed 2024).
  25. Iverson, J. B. 1991c. Patterns of survivorship in turtles (order Testudines). Canadian J. Zoology 69:385-391.
  26. Iverson, J. B., and G. R. Smith. 1993. Reproductive ecology of the painted turtle (<i>Chrysemys picta</i>) in the Nebraska Sandhills and across its range. Copeia 1993:1-21.
  27. Jensen, E.L., P. Govindarajulu, and M.A. Russello. 2014. When the shoe doesn’t fit: applying conservation unit concepts to western painted turtles at their northern periphery. Conservation Genetics 15(2):261-274.
  28. Jensen, E.L., P. Govindarajulu, and M.A. Russello. 2015. Genetic assessment of taxonomic uncertainty in painted turtles. Journal of Herpetology 49(2):314-324.
  29. King, F. W., and R. L. Burke, editors. 1989. Crocodilian, tuatara, and turtle species of the world: a taxonomic and geographic reference. Association of Systematics Collections, Washington, D.C. 216 pp.
  30. Laporte, M., C.-O. Silva Beaudry, and B. Angers. 2012. Effects of road proximity on genetic diversity and reproductive success of the painted turtle (<i>Chrysemys picta</i>). Conservation Genetics 14: 21-30.
  31. Lindeman, P. V. 1991. Survivorship of overwintering hatchling painted turtles, <i>Chrysemys picta</i>, in northern Idaho. Can. Field-Nat. 105:263-266.
  32. Lindeman, P. V. 1996. Comparative life history of painted turtles (<i>Chrysemys picta</i>) in two habitats in the inland Pacific Northwest. Copeia 1996:114-130.
  33. Lindeman, P. V. 2014. New wine in old bottles: using modified hoopnets to catch bait-averse basking turtles. Herpetological Review 45(4): 597-600.
  34. Lindeman, P. V., and A. F. Scott. 2001. Over three decades of persistence of a small and apparently isolated population of painted turtles (Chrysemys picta) in a Kentucky reservoir. Chelonian Conservation and Biology 4:206-208.
  35. MacCulloch, R. D., and D. M. Secoy. 1983. Movement in a river population of <i>Chrysemys picta belli</i> in southern Saskatchewan. Journal of Herpetology 17:283-285.
  36. Marchand, K. A., R. G. Poulin, and C. M. Somers. 2018. Western painted turtles (<i>Chrysemys picta bellii</i>) in a highly urbanized system: unexpected variation in resource use among age classes and sexes. Herpetologica 74(3): 217-225.
  37. McAuliffe, J. R. 1978. Seasonal migrational movements of a population of the western painted turtle, <i>Chrysemys picta bellii</i> (Reptilia, Testudines, Testudinidae). Journal of Herpetology 12:143-149.
  38. Mitchell, J. C. 1988. Population ecology and life histories of the freshwater turtles <i>Chrysemys picta</i> and <i>Sternotherus odoratus</i> in an urban lake. Herpetol. Monogr. 2:40-61.
  39. Moll, E. O. 1973. Latitudinal and intersubspecific variation in reproduction of the painted turtle, <i>Chrysemys picta</i>. Herpetologica 29:307-318.
  40. Packard, G. C., and M. J. Packard. 1995. A review of the adaptive strategy used by hatchling painted turtles to survive in the cold. Herpetological Review 26:8, 10-12.
  41. Rowe, J. W. 2003. Activity and movements of midland painted turtles (<i>Chrysemys picta marginata</i>) living in a small marsh system on Beaver Island, Michigan. Journal of Herpetology 37:342-353.
  42. Starkey, D. E., H. B. Shaffer, R. L. Burke, M.R.J. Forstner, J. B. Iverson, F. J. Janzen, A.G.J. Rhodin, and G. R. Ultsch. 2003. Molecular systematics, phylogeography, and the effects of Pleistocene glaciation in the painted turtle (<i>Chrysemys picta</i>) complex. Evolution 57:119-128.
  43. Stebbins, R. C. 2003. A field guide to western reptiles and amphibians. Third edition. Houghton Mifflin Company, Boston.
  44. Turtle Taxonomy Working Group (TTWG) [Rhodin, A. G. J., J. B. Iverson, R. Bour, U. Fritz, A. Georges, H. B. Shaffer, and P. P. van Dijk]. 2021. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9th Ed.). In: Rhodin, A. G. J., J. B. Iverson , P. P. van Dijk, C. B. Stanford, E. V. Goode, K. A. Buhlmann, and R. A. Mittermeier (Eds.). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs 8:1–472. doi: 10.3854/crm.8.checklist.atlas.v9.2021.
  45. Turtle Taxonomy Working Group [van Dijk, P.P., Iverson, J.B., Shaffer, H.B., Bour, R., and Rhodin, A.G.J.]. 2012. Turtles of the world, 2012 update: annotated checklist of taxonomy, synonymy, distribution, and conservation status. In: Rhodin, A.G.J., Pritchard, P.C.H., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Iverson, J.B., and Mittermeier, R.A. (Eds.). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs No. 5:000.243-000.328. Online. Available: www.iucn-tftsg.org/cbftt/.
  46. Ultsch, G. R., G. M. Ward, C. M. LeBerte, B. R. Kuhajda, and E. R. Stewart. 2001. Intergradation and origins of subspecies of the turtle Chrysemys picta: morphological comparisons. Canadian Journal of Zoology 79:485-498.
  47. Vogt, R. C. 1981c. Natural history of amphibians and reptiles of Wisconsin. Milwaukee Public Museum. 205 pp.
  48. Washington Herp Atlas. 2009 (map products updated March 2017). A cooperative effort of Washington Natural Heritage Program, Washington Department of Fish and Wildlife, U.S.D.I. Bureau of Land Management, and U.S. Forest Service. 250 pp.